bims-traimu Biomed News
on Trained immunity
Issue of 2026–02–15
eight papers selected by
Yantong Wan, Southern Medical University



  1. Sci Rep. 2026 Feb 10.
      We previously demonstrated that Bacillus Calmette Guérin (BCG) vaccination induces trained immunity in bovine monocytes and gamma delta (γδ) T cells, enhancing IL-1β and IL-6 production. To define the underlying molecular mechanisms, we applied a multi-omic approach to circulating monocytes and alveolar macrophages (amac). Twenty Holstein-Angus heifers were randomly separated into two groups and immunized subcutaneously with 6 million CFUs of BCG Danish strain (n = 10), or saline (n = 10), and boosted after 2 weeks. Monocytes (4 weeks post-boost) and amac (6 weeks post-boost) were analyzed for cytokine production, transcriptional responses (QuantSeq 3' mRNA sequencing), and histone modifications (CUT&Tag H3K27ac). BCG vaccination increased IL-1β (p = 0.020 and IL-6 (p < 0.0001) production in monocytes, and increased IL-1β (p = 0.041) in amac following heterologous stimulation. Transcriptomic analysis revealed an expanded breadth of stimulation-responsive genes in BCG-trained monocytes, (1439 vs 1214 differentially expressed genes (DEGs, FDR < 0.05) with LPS; 6513 vs 5605 DEGs with Poly(I:C)/Imiquimod). Gene ontology analysis demonstrated enrichment of innate immune response pathways among BCG-enhanced genes. H3K27ac profiling identified 227 differential peaks (DP, p < 0.01, |FC|> 1.25) in monocytes, most of which are bovine-specific compared to human, and 383 DP in amac.These datasets provide evidence that BCG induces trained immunity in bovine monocytes and macrophages through enhanced cytokine capacity, transcriptional reprogramming, and epigenetic modifications.
    Keywords:  BCG vaccination; Bovine; Epigenetics; H3K27ac; Trained immunity; Transcriptomics
    DOI:  https://doi.org/10.1038/s41598-026-39580-7
  2. JCI Insight. 2026 Feb 10. pii: e195866. [Epub ahead of print]
      Infectious diseases remain a global health challenge, driven by increasing antimicrobial-resistance and the threat of emerging epidemics. Mycobacterium tuberculosis and Staphylococcus aureus are leading causes of mortality worldwide. Trained immunity-a form of innate immune memory-offers a promising approach to enhance pathogen clearance. Here, we demonstrate that IFN-γ induces trained immunity in human monocytes through a mechanism involving mTORC1 activation, glutaminolysis, and epigenetic remodeling. Macrophages derived from IFN-γ-trained monocytes exhibited increased glycolytic activity with enhanced cytokine and chemokine responses upon stimulation or infection. Crucially, trained macrophages had increased production of reactive oxygen species which mediated enhanced bactericidal activity against methicillin-resistant S. aureus. Furthermore, ATAC-sequencing analysis of IFN-γ trained macrophages revealed increased chromatin accessibility in regions associated with host defence. Lastly, IFN-γ training restored impaired innate responses in macrophages from individuals homozygous for the TIRAP 180L polymorphism, a genetic variant associated with increased susceptibility to infection. These findings establish IFN-γ as a potent inducer of trained immunity in human monocytes and support its potential as a host-directed strategy to strengthen antimicrobial defenses, particularly in genetically susceptible individuals and high-risk clinical contexts.
    Keywords:  Bacterial infections; Immunology; Infectious disease; Macrophages; Preventative medicine
    DOI:  https://doi.org/10.1172/jci.insight.195866
  3. Biology (Basel). 2026 Jan 23. pii: 213. [Epub ahead of print]15(3):
      Hypoxia is a common feature of inflamed and ischemic tissues and represents an important regulatory signal for innate immune cells. The master regulator of this response is hypoxia-inducible factor-1α (HIF-1α), a transcription factor whose stabilization and activity are tightly regulated by the presence of oxygen, inflammatory signaling, and cellular metabolism. Monocytes, key players in innate immunity, rapidly sense oxygen deprivation and display specific responses during acute hypoxia, primarily aimed at adapting and maintaining cellular homeostasis. Unlike macrophages, in which HIF-1α activity is known, the mechanisms regulating HIF-1α stabilization, subcellular localization, and transcriptional activity in circulating monocytes remain incompletely elucidated. Recent studies indicate that acute hypoxia primarily triggers post-translational stabilization of HIF-1α, calcium- and PKC-dependent signaling, metabolic reprogramming, and early inflammatory responses, while transcriptional activation of HIF-1α may require additional inflammatory or stress-related signals. Furthermore, extensive crosstalk between HIF-1α and NF-κB integrates hypoxic and inflammatory signals, modulating cytokine production, cell migration, and survival. Epigenetic regulators can also modulate these responses and contribute to hypoxia-induced trained immunity. In this review, we summarize current knowledge of the mechanisms controlling the stabilization, localization, and function of HIF-1α in human monocytes and monocyte-macrophages during acute hypoxia, highlighting the key differences between these cell types and discussing their implications for inflammation, tissue homeostasis, and disease.
    Keywords:  HIF-1α; NF-κB; cytokines; inflammation; innate immunity; macrophages
    DOI:  https://doi.org/10.3390/biology15030213
  4. Front Med (Lausanne). 2026 ;13 1721780
       Introduction: Chronic obstructive pulmonary disease (COPD) is a heterogeneous inflammatory disorder characterized by persistent immune dysregulation and progressive structural deterioration of the lung. However, how COPD reshapes lung architecture, immune signaling, and cellular identity at a systems level remains incompletely understood.
    Methods: We performed integrative, multi-dimensional transcriptomic analysis of human COPD lung datasets to evaluate alterations in immune signaling, regulated cell death pathways, fibrosis-associated programs, cell type-specific transcriptional identity, and immune checkpoint regulation. Genetic and cytokine-based perturbations targeting trained immunity pathways were analyzed to assess functional relevance.
    Results: COPD induced broad transcriptional activation of cytokines, secretory and plasma membrane proteins, CD markers, innate immune genes, and trained immunity genes. Deficiency of SET7, a promoter of trained immunity, or overexpression of IL-37, an inhibitor of trained immunity, attenuated expression of COPD-upregulated immune genes. COPD also promoted tissue injury through coordinated upregulation of genes regulating multiple forms of regulated cell death, including autosis, autophagy, parthanatos, immunogenic cell death, mitochondrial permeability transition-associated death, lysosomal cell death, apoptosis, necroptosis, ferroptosis, mitotic cell death, and proliferation-associated cell death. In parallel, COPD enhanced epithelial-to-mesenchymal transition and fibrosis-related transcriptional programs. Transcriptomic identity was disrupted in 10 of 14 major human lung cell types, with evidence of pathological trans-differentiation marked by aberrant expression of over 50 cell type-specific marker genes. Alveolar macrophages exhibited extensive dysregulation of immune checkpoint ligand; notably, PVR (CD155) expression was reduced in severe emphysema, while experimental PVR overexpression suppressed pro-inflammatory gene expression in both alveolar and interstitial macrophages. Additionally, COPD impaired the suppressive capacity of CD4+Foxp3+ regulatory T cells through downregulation of key immunosuppressive genes, including those associated with FoxP3+, TIGIT+, and LPS-responsive Tregs. Shared immunosuppressive gene signatures were identified between PVR-overexpression-inducing CD4+ T cells and IL-10-mediated regulatory pathways in T cells and monocytes.
    Discussion: Collectively, these findings demonstrate that COPD reprograms the lung toward an immune-like organ by promoting immune cell-like trans-differentiation of structural cells, activating diverse regulated cell death pathways, and altering immune checkpoint signaling. These mechanisms highlight potential therapeutic targets for immunomodulatory intervention in COPD.
    Keywords:  COPD; epithelial-to-mesenchymal transition (EMT); immune organs; secretomes; trained immunity
    DOI:  https://doi.org/10.3389/fmed.2026.1721780
  5. Hum Vaccin Immunother. 2026 Dec;22(1): 2624234
      Aging reshapes immunity through immunosenescence and inflammaging, increasing susceptibility to infection, exacerbating chronic conditions, and blunting vaccine responses. This review frames "immunofitness" as a practical goal of healthy aging and examines how adult vaccination builds immune resilience. Vaccination strengthens adaptive memory, leverages adjuvants to optimize antigen presentation, and can reprogramme innate cells (trained immunity), yielding heterologous benefits beyond target pathogens. We integrate evidence in older adults for influenza, respiratory syncytial virus, pneumococcal, COVID-19, and recombinant zoster vaccines, including reductions in respiratory events, cardiovascular outcomes, hospitalization, and mortality. We highlight emerging platforms and precision vaccinology to tailor schedules by immune age, comorbidity, and frailty. Integrating routine, age-appropriate vaccination with lifestyle measures is a feasible, high-impact strategy to promote immunofitness.
    Keywords:  Immunofitness; adult vaccination; immunosenescence; inflammaging; older adults; trained immunity
    DOI:  https://doi.org/10.1080/21645515.2026.2624234
  6. Commun Biol. 2026 Feb 12. 9(1): 239
      Interferon regulatory factors (IRFs) are essential for transcription of interferons (IFNs), interferon-stimulated genes (ISGs), and pro-inflammatory cytokines. We profile the transcriptome of human monocyte THP1 cells challenged with cGAMP, LPS, or IFNB1 protein as a function of knockout (KO) or overexpression (OE) of IRFs or KO of IFNAR2. We define distinct gene expression groups, reflecting the transcription factors responsible for their induction including subgroups activated by more than one pathway or feed-forward regulation. We compare IRF3- and IRF7-induced gene signatures and note the strong direct induction of a subset of antiviral-acting ISGs by IRF3 or IRF7. LPS treatment induces NF-κB responses in monocyte and macrophage state cells, however, IFNs and ISGs are only co-induced in the macrophage state requiring IRF3. IRF1, IRF2, IRF5, and IRF8 are largely dispensable for IFN-stimulated or innate-immune-mediated gene induction. This study provides a valuable resource for dissecting complex inflammatory gene signatures and their underlying transcription factors thereby anticipating the effects of selectively drugging the underlying pathways.
    DOI:  https://doi.org/10.1038/s42003-025-09343-7
  7. Cell Rep. 2026 Feb 10. pii: S2211-1247(26)00063-X. [Epub ahead of print]45(2): 116985
      Interferon γ (IFN-γ) is a key cytokine in immune activation, especially anti-viral responses, and a driver of macrophage activation. It classically signals via JAK1/2-mediated STAT1 homodimers. Here, we identify an alternative, non-canonical signaling component in which IFN-γ simultaneously also activates STAT3. Our results show that IFN-γ activates STAT3 rapidly and directly through JAK1 and JAK2. We provide evidence that STAT3 can form heterodimers with STAT1 in this context and demonstrate that STAT3 is co-recruited to a subset of IFN-γ-induced, STAT1-bound regulatory elements. While IFN-γ directly activates STAT3, our results reveal that its contribution to IFN-γ-induced gene expression is limited. Instead, our data indicate that STAT1-STAT3 heterodimers exert a more prominent function in mixed cytokine environments. These findings uncover STAT3 as an unconventional player in macrophage IFN-γ signaling, underscoring the complex and context-dependent nature of cytokine signaling networks.
    Keywords:  CP: immunology; IFN-γ; JAK; STAT; cytokine; interferon; macrophages; transcription
    DOI:  https://doi.org/10.1016/j.celrep.2026.116985
  8. Phytomedicine. 2026 Jan 28. pii: S0944-7113(26)00133-9. [Epub ahead of print]153 157894
       BACKGROUND: Excessive inflammatory responses mediated by innate immunity are a significant cause of sepsis-induced acute lung injury (SI-ALI), underscoring the need for further drug development to improve therapeutic outcomes. Ginkgetin (GK), a natural flavone, has demonstrated potential anti-inflammatory and antioxidative effects. However, it remains unclear whether GK can ameliorate SI-ALI and what the underlying mechanisms might be.
    PURPOSE: We aimed to investigate the therapeutic effects and mechanisms of GK on SI-ALI.
    STUDY DESIGN/METHODS: We established SI-ALI models using lipopolysaccharide or cecal ligation and puncture to assess the impact of GK. An autophagy agonist and inhibitor were applied to detect the effect of GK on autophagy. Proteomics and targeted gene knockdown experiments were employed to validate lysosomal-associated transmembrane protein 5 (Laptm5) as the key protein. Co-immunoprecipitation and site mutation assays were used to identify the site where GK inhibited Laptm5 ubiquitination. Biotin pulldown coupled with mass spectroscopy, molecular docking, and drug affinity responsive target stability (DARTS) were performed to elucidate the direct target and underlying mechanisms.
    RESULTS: We demonstrated that GK activated the autophagosome-lysosome pathway by increasing the protein level of Laptm5. This process facilitated autophagy-mediated degradation of tank-binding kinase 1 (TBK1) and inhibited the signal transduction of downstream inflammatory pathways. Further, we found that GK inhibited the K48-linked ubiquitination of Laptm5 and revealed the ubiquitination sites of Laptm5 (K86 and K122) for the first time. Biotin pulldown and DARTS identified ubiquitin-protein ligase E3C (Ube3c) as a target of GK in inhibiting Laptm5 ubiquitination, with TYR707 and ASN832 being the key residues.
    CONCLUSION: Our findings indicate that GK exerts its anti-inflammatory effect on macrophages by promoting autophagy via suppressing the ubiquitination of Laptm5, thus offering a promising therapeutic approach for SI-ALI.
    Keywords:  Acute lung injury; Autophagy; Ginkgetin; Laptm5; Macrophage; Ubiquitination
    DOI:  https://doi.org/10.1016/j.phymed.2026.157894