bims-pideca Biomed News
on Class IA PI3K signalling in development and cancer
Issue of 2026–02–22
seventeen papers selected by
Ralitsa Radostinova Madsen, MRC-PPU



  1. Mol Cell Proteomics. 2026 Feb 17. pii: S1535-9476(26)00026-5. [Epub ahead of print] 101530
      The increasing scale and complexity of proteomics data demand robust, scalable, and interpretable quality control (QC) frameworks to ensure data reliability and reproducibility. Here, we present pmultiqc, an open-source Python package that standardizes and generates web-based QC reports across multiple proteomics data analysis platforms. Built on top of the widely adopted MultiQC framework, pmultiqc offers specialized modules tailored to mass spectrometry workflows, with full initial support for quantms, DIA-NN, MaxQuant/MaxDIA, FragPipe, and mzIdentML/mzML-based pipelines. The package computes a wide range of QC metrics, including raw intensity distributions, identification rates, retention time consistency, and missing value patterns, and presents them in interactive, publication-ready reports. By leveraging sample metadata in the SDRF format, pmultiqc enables metadata-aware QC and introduces, for the first time in proteomics, QC reports and metrics guided by standardized sample metadata. Its modular architecture allows easy extension to new workflows and formats. Alongside comprehensive documentation and examples for running pmultiqc locally or integrated into existing workflows, we offer a cloud-based service that enables users to generate QC reports from their own data or public PRIDE datasets.
    DOI:  https://doi.org/10.1016/j.mcpro.2026.101530
  2. Cell J. 2025 Dec 22. pii: 731598. [Epub ahead of print]27(1): 1-12
       Objective: The PI3K/Akt signaling pathway plays a central role in regulating cell growth, survival, and metabolism, and its dysregulation is a hallmark of many cancers. The PIK3CA gene, which encodes the alpha catalytic subunit of PI3K, is altered in approximately 30% of breast cancers. Among its mutations, c.3140A>G (p.His1047Arg) in the kinase domain is the most prevalent, producing a constitutively active enzyme with oncogenic potential. Here, we engineered a population of MCF7 cells carrying the PIK3CA c.3140A>G mutation using CRISPR-Cas9 with precise single-nucleotide editing, and evaluated its impact on cellular characteristics.
    Materials and Methods: In this experimental study, nearly homogeneous populations of PIK3CA H1047R mutant MCF7 cells were generated using CRISPR-Cas9-mediated genome editing followed by hierarchical single-cell isolation. Editing efficiency was validated through allele-specific polymerase chain reaction (PCR) and multiple rounds of Sanger sequencing. Cell cycle distribution and proliferation were analyzed using flow cytometry and cell count assays, respectively. Gene expression changes were assessed by quantitative real-time PCR to evaluate the mutation's impact on cell cycle-related genes.
    Results: Tracking of insertions, deletions, and recombination events (TIDER) analysis showed approximately 60% homology-directed repair (HDR) efficiency in the edited population. Flow cytometry revealed a 5% increase in the G2/M cell population in the edited clone compared with unedited controls (P<0.001). Proliferation assays demonstrated significantly accelerated growth (1.30 fold) under low fetal bovine serum (FBS) conditions (P=0.029). Quantitative real-time PCR confirmed upregulation of cell cycle-promoting genes, with CCND1 and MYC expression increasing by 1.62-fold (P<0.001) and 1.23-fold (P<0.001), respectively, relative to controls.
    Conclusion: The genetically edited cell lines represent robust and well-defined experimental models that enable direct assessment of the functional consequences of oncogenic driver mutations on cellular behavior and signaling pathways. Our findings demonstrate that targeted genetic alterations induce measurable changes in proliferation, cell-cycle regulation, and gene expression, thereby providing mechanistic insight into tumorigenesis and the specific contribution of driver mutations to cancer-related cellular phenotypes.
    Keywords:  Breast Cancer; CRISPR-Cas9; Genome EditingGenome Editing; MCF7 Cell Line; PIK3CA
    DOI:  https://doi.org/10.22074/cellj.2025.2055578.1813
  3. J Exp Med. 2026 Apr 06. pii: e20252154. [Epub ahead of print]223(4):
      While inputs regulating CD4+ T helper (Th) cell differentiation are well defined, the integration of downstream signaling with transcriptional and epigenetic programs that define Th lineage identity remains incompletely resolved. PI3K signaling is a critical regulator of T cell function; activating mutations affecting PI3Kδ result in an immunodeficiency with multiple T cell defects. Using mice expressing activated PI3Kδ, we found aberrant expression of proinflammatory Th1 signature genes under Th2-inducing conditions, both in vivo and in vitro. This dysregulation was driven by a PI3Kδ-IL-2-Foxo1 signaling amplification loop, fueling Foxo1 inactivation, loss of Th2 lineage restriction, and extensive epigenetic reprogramming. Surprisingly, ablation of Fasl, a Foxo1-repressed gene, normalized both Th2 differentiation and TCR signaling. BioID and imaging revealed Fas interactions with TCR signaling components, which were supported by Fas-mediated potentiation of TCR signaling that could occur in the absence of FADD. Our results highlight Fas-FasL signaling as a critical intermediate in phenotypes driven by activated PI3Kδ, thereby linking two key pathways of immune dysregulation.
    DOI:  https://doi.org/10.1084/jem.20252154
  4. J Clin Invest. 2026 Feb 16. pii: e194300. [Epub ahead of print]136(4):
      Germline loss-of-function folliculin (FLCN) gene mutations cause Birt-Hogg-Dubé (BHD) syndrome, in which pulmonary cysts are present in up to 90% of the patients. The pathogenic mechanisms underlying lung cyst development in BHD are almost entirely unknown because of the limited availability of BHD patient lung samples and the lack of authentic BHD lung disease models. We generated lung mesenchyme-specific and lung epithelium-specific Flcn-knockout mice using a Cre/loxP approach. We found that deletion of Flcn in lung mesenchymal cells, but not in lung epithelial cells, resulted in alveolar enlargement starting from early postnatal life, with evidence of cyst formation in adult mice, resembling the pulmonary disease in human BHD. These changes were associated with increased mechanistic target of rapamycin complex 1 (mTORC1) activity in the lungs of both patients with BHD and Flcn-knockout mice. Attenuation of mTORC1 activity by knocking out Raptor gene (Rptor) or pharmacologic inhibition using rapamycin substantially rescued the pulmonary pathology caused by Flcn deletion in mice. Taken together, these human and mouse data support a model in which mTORC1 hyperactivation drives pulmonary cystic pathology in BHD.
    Keywords:  Cell biology; Development; Mouse models; Pulmonology; Tumor suppressors
    DOI:  https://doi.org/10.1172/JCI194300
  5. Nat Rev Mol Cell Biol. 2026 Feb 18.
      Biological functions depend on the spatiotemporal distribution of proteins within cells. Key cellular activities such as signal transduction, metabolism, cell cycle and cell death are driven by the interactions of proteins that are localized in multiple cellular compartments. Such multilocalization can even allow protein with identical sequences to display multifunctionality, a phenomenon known as moonlighting. Despite its biological importance, the relationship between protein localization and function remains underexplored. In this Review, we discuss the known mechanisms of protein localization (including RNA transport, role of proteoforms and molecular interactions) and how subcellular localization controls protein function. Proper regulation of protein localization is crucial for specialized cell and tissue functions, including cell differentiation, polarization and the epithelial-mesenchymal transition. Protein mislocalization can also have important roles in pathological processes, such as in cancer, neurodegeneration and autoimmunity. We end with a discussion of current technological and conceptual challenges in the field of subcellular proteomics and spatial biology. Addressing these challenges will allow us to link the dynamic nature of protein localization and function across biological scales and contexts, with great impact on fundamental cell biology and clinical applications.
    DOI:  https://doi.org/10.1038/s41580-026-00947-3
  6. Development. 2026 Feb 15. pii: dev205542. [Epub ahead of print]153(4):
      Soumyashree Das is an Assistant Professor at the National Centre for Biological Sciences at the Tata Institute of Fundamental Research in Bangalore, India. Her research focuses on endothelial cell fate decisions in collateral artery formation, using a range of mouse models, imaging approaches, and other cellular and molecular techniques. The goal of her lab is to understand how these arteries form in different tissues. We spoke to Soumyashree over Zoom to find out more about her career path, her research on vascular development and regeneration, and her experience establishing a new lab under pandemic conditions.
    DOI:  https://doi.org/10.1242/dev.205542
  7. EMBO Mol Med. 2026 Feb 18.
      Somatic activating KRAS mutations in endothelial cells are the predominant cause of sporadic brain arteriovenous malformations (bAVMs) and also occur in sporadic extracranial AVMs. We found that KRASG12V expression in the endothelium increased angiogenesis, which was accompanied by enhanced glucose uptake and glycolytic flux. Mechanistically, this increase in glycolysis was facilitated by enhanced membrane localization of glucose transporters (e.g., GLUT1) and induction of hexokinase-2 (HK2) expression. Importantly, RNA-sequencing and proteomics revealed that HK2 appeared to be the only glycolytic component elevated. Analysis of single-cell RNA-sequencing data and immunofluorescence staining confirmed that HK2 was elevated in mouse and human bAVMs. Critically, either pharmacologic inhibition of glycolytic flux or knockdown of HK2 suppressed sprouting angiogenesis in cultured KRASG12V endothelial cells. Glycolysis inhibition also reversed arteriovenous shunts and potentiated the effect of MEK inhibition in a KRAS-mutant zebrafish model. Finally, combined glycolysis and MEK inhibition suppressed angiogenesis in patient-derived bAVM primary endothelial cells. Together, our findings show that KRAS-driven reprogramming of endothelial metabolism represents a potential therapeutic vulnerability for sporadic AVMs.
    Keywords:  Arteriovenous Malformations; Endothelial Glycolysis; KRAS Signaling; Metabolic Regulation; Targeted Therapeutics
    DOI:  https://doi.org/10.1038/s44321-026-00383-y
  8. Cell Syst. 2026 Feb 18. pii: S2405-4712(26)00016-5. [Epub ahead of print]17(2): 101534
      Deriving principles governing cell biology from single-cell measurements across modalities, called multimodal modeling, can advance our understanding of cellular states in health and disease. Realizing the full potential of multimodal models requires learning generalizable representations across data types, diseases, and biological contexts. This perspective examines the potential of compositional AI as a modular design approach for constructing multimodal foundation models that unify biological modalities-such as chromatin accessibility, protein abundance, spatial transcriptomics, microscopy imaging, and textual annotations-into cohesive representations of cellular behavior. We present key deep learning modeling approaches, along with transformer-based attention strategies to implement them, while addressing challenges posed by limited data availability and structural differences between modality representations. We also discuss how to connect and align partially overlapping multimodal measurements to build a comprehensive representation space. By synthesizing these technical advancements, we chart a path toward agentic virtual cell models, offering insights into opportunities, limitations, and future directions for leveraging multimodal AI to decode the complexity of cellular systems.
    Keywords:  compositional AI; single-cell foundation models; single-cell multi-omics
    DOI:  https://doi.org/10.1016/j.cels.2026.101534
  9. Methods Enzymol. 2026 ;pii: S0076-6879(25)00520-8. [Epub ahead of print]726 45-83
      Cells derived from diseased tissue and their related cell lines exhibit numerous metabolic changes, including variations in lipid composition and metabolism. Indeed, lipids are important biomarkers of various diseases and exhibit crucial signaling roles during disease states. However, lipids, especially low-abundant and transient lipids like phosphoinositides, are difficult to study due to a lack of sophisticated tools. Here, we describe a unique targeted lipidomics method that allows us to define and compare the phosphoinositide composition of diseased and healthy tissues as well as related cell lines.
    Keywords:  ESI-MS/MS; Lipid analysis; Lipidomics; Phosphatidylinositol; Phosphatidylinositol phosphates; Phosphatidylinositol trisphosphate; Phosphoinositides; Targeted mass spectrometry
    DOI:  https://doi.org/10.1016/bs.mie.2025.11.016
  10. Adv Sci (Weinh). 2026 Feb 19. e16423
      Epithelial and endothelial monolayers maintain homeostasis by adapting to physiological stimuli and injury through conversion processes that remain incompletely understood. Using human umbilical vein endothelial cell cultures (HUVECs), we elucidate how monolayer maturation and mechanotransduction-induced remodeling are molecularly regulated. Maturation involves reduced cell perimeter leading to increased junctional VE-cadherin that recruits junctional actin, integrins and vinculin to establish a quiescent, stable monolayer. Remarkably, we identify a previously unrecognized, rapid and reversible intermediate-state, marked by VE-cadherin linearization (clustering) and actomyosin relaxation via myosin light chain (MLC)-dephosphorylation, that emerges during mechanotransduction-induced activation, triggered by onset or shifts in shear stress-induced mechanical load. This novel tension-mediated intermediate state enhances junctional actin, integrin and vinculin recruitment, thereby strengthening barrier function while protecting endothelial cells from overstimulation and mechanical damage. MLC rephosphorylation dissolves junctional actin, forms stress fibers and induces the formation of "Junction-Associated-Intermittent-Lamellipodia" (JAIL), enabling cell shape change and arterial phenotype remodeling. Overall, junctional VE-cadherin concentration, together with mechanosensitive signaling that reduces actomyosin tension, governs actin recruitment, revealing a tension-sensitive, intermediate state that protects cells and primes endothelial remodeling. The data provide a broader model for endothelial mechanotransduction and stress adaptation.
    Keywords:  actin‐dynamics; actomyosin contractility; cell junction dynamics; cellular conversion; integrin dynamics; mechanical stress; mechanotransduction
    DOI:  https://doi.org/10.1002/advs.202516423
  11. J Biol Chem. 2026 Feb 12. pii: S0021-9258(26)00152-3. [Epub ahead of print] 111282
      The plasma membrane (PM) is a dynamic interface that integrates environmental cues with cellular responses. Insulin is known to remodel the PM primarily by stimulating the translocation of glucose transporter GLUT4, but the full scope of insulin's PM remodeling remains poorly defined. Here, we performed a meta-analysis of insulin-regulated PM proteins in adipocytes by integrating nine independent proteomic datasets generated using complementary PM enrichment strategies. The meta-analysis identified 37 insulin-regulated candidates detected in at least three datasets, including 30 proteins not previously implicated in insulin action. Among these, we experimentally characterized the insulin-stimulated translocation of two transporters: potassium-chloride cotransporter 1 KCC1 (SLC12A4) and sodium-dependent phosphate transporter PIT2 (SLC20A2), which showed robust and reproducible recruitment to the PM in response to insulin. siRNA-mediated knockdown of KCC1 or PIT2 impaired insulin-stimulated glucose transport, suggesting a role for these transporters in insulin action. Live-cell and fixed-cell imaging revealed that both proteins localize across multiple endosomal compartments, undergo insulin dose-dependent trafficking to the PM, and require PI3K-AKT signaling for their mobilization. Strikingly, insulin-induced translocation of KCC1 and PIT2 to the PM was impaired in adipocytes rendered insulin resistant by chronic hyperinsulinemia, accompanied by increased perinuclear retention under basal conditions. Together, our work provides a valuable resource for understanding insulin-regulated PM remodeling in adipocytes, establishes KCC1 and PIT2 as novel insulin-responsive transporters, and supports the idea that insulin resistance involves defects in cell-surface delivery that extend beyond GLUT4.
    Keywords:  GLUT4; KCC1; PIT2; adipocyte; insulin; insulin resistance; plasma membrane; proteomics; trafficking
    DOI:  https://doi.org/10.1016/j.jbc.2026.111282
  12. iScience. 2026 Feb 20. 29(2): 114784
      Tumor vasculature is abnormally formed, with an endothelial cell (EC) barrier lacking integrity, resulting in hyperpermeable vessels. Elevated VEGFA levels drive the formation of the dysfunctional vasculature by activating VEGFR2 signaling. The VEGFR2 tyrosine site pY1175 was recently shown to stimulate a PLCγ/eNOS/Src pathway, promoting vascular leakage and hindering therapeutic targeting and anti-tumor immunity. High endothelial PLCγ levels correlated to poor kidney cancer prognosis, which indicated endothelial PLCγ as a prognostic biomarker. In this study, we reveal SHP2 as a binding partner of pY1175 and show that SHP2 cooperates with PLCγ to mediate VEGFA-induced permeability in both healthy and tumor vasculatures. Targeting the VEGFR2/PLCγ/SHP2 axis-genetically or pharmacologically-reduces EC junctional phosphorylation to prevent VE-cadherin internalization, followed by reduced macromolecular leakage. Tumor EC expression of PLCγ or SHP2 is associated with vascular leakage in human kidney cancer, underscoring their potential as targets for vascular normalization and biomarkers for disease progression and treatment response.
    Keywords:  cancer; cell biology; microenvironment; molecular biology
    DOI:  https://doi.org/10.1016/j.isci.2026.114784
  13. Cell Stem Cell. 2026 Feb 13. pii: S1934-5909(26)00030-5. [Epub ahead of print]
      Transgenes are often silenced upon differentiation of pluripotent stem cells using conventional expression systems. Here, we developed the TK4 PiggyBac vector to conduct a comparative analysis to evaluate the impact of various promoters, transcriptional regulatory elements, insulators, and genomic integration sites on transgene silencing during neuronal differentiation. Our findings reveal that specific combinations of CAG and Ubc promoters with the Woodchuck hepatitis virus post-transcriptional regulatory element (WPRE) can prevent transgene silencing during differentiation, whereas chromatin insulators have less impact on sustained expression. Three novel safe harbor loci, distant from known genes, as well as the citrate lyase beta-like (CLYBL) locus, similarly support the prevention of transgene silencing. Remarkably, the TK4 vector showed complete resistance to silencing across various neuronal and microglial differentiation protocols, as independently confirmed by seven laboratories. This construct will be highly useful for assays requiring stable transgene expression during differentiation and holds potential for broad applications in various research fields.
    Keywords:  PiggyBac vector; UCOE; WPRE; chromatin insulator; human pluripotent stem cells; microglia; neuron; promoter activity; safe harbor locus; transgene silencing; ubiquitous chromatin opening element; woodchuck hepatitis posttranscriptional regulatory element
    DOI:  https://doi.org/10.1016/j.stem.2026.01.007
  14. Cell Syst. 2026 Feb 18. pii: S2405-4712(26)00020-7. [Epub ahead of print]17(2): 101538
      Biomedical research requires quantitative rigor, i.e., numeracy, a facility with numbers. The last decade has seen the broad adoption of statistical tools ("Numeracy 1.0"). To drive science forward, the expertise to quantitatively evaluate hypotheses and insights also needs to be broadly adopted ("Numeracy 2.0"). Systems biologists will be at the forefront of the transformation.
    DOI:  https://doi.org/10.1016/j.cels.2026.101538
  15. Cell Metab. 2026 Feb 17. pii: S1550-4131(26)00012-4. [Epub ahead of print]
      Endothelial cells (ECs) are key metabolic gatekeepers, yet their role in metabolic health remains unclear. Given their central involvement in energy metabolism, mitochondria are ideally positioned to enable ECs to adapt to ever-changing metabolic requirements. Here, we explore the hypothesis that mitochondrial dynamics proteins in ECs influence whole-body metabolic status. Genetic deficiency of Mfn2 in ECs (Mfn2iΔEC), but not of Mfn1iΔEC, induces a mitohormetic response in the adipose vasculature, enhancing antioxidant defenses, mitochondrial fitness, and lipid oxidation, ultimately improving metabolic outcomes. Cultured ECs secrete the mitokine growth differentiation factor 15 (GDF15) via a forkhead box O1 (FOXO1)-dependent axis, a response also observed under stress conditions in vivo. Notably, Mfn2iΔEC mice exhibited elevated endothelial and circulating GDF15 levels, and neutralization of GDF15 partly attenuated their metabolic benefits. Consistent with mitohormetic activation, Mfn2iΔEC mice showed protection against diet-induced obesity and delayed age-related decline. Hence, vascular mitohormetic adaptations emerge as a novel mechanism promoting systemic metabolic health.
    Keywords:  GDF15; aging; diabetes; endothelial cells; mitochondria; mitofusin; mitohormesis; obesity
    DOI:  https://doi.org/10.1016/j.cmet.2026.01.012
  16. Nat Mater. 2026 Feb 20.
      Increased extracellular matrix deposition and stiffness promotes solid tumour progression. Yet, the precise mechanotransduction pathways, especially in less-studied mechanically responsive cancers, remain poorly understood. Here we address this gap using patient-derived tumour cells from early (mobile, T1) and advanced (immobile, T3) stages of vocal fold cancer, the most common squamous cell carcinoma severely impacting the voice box. We reveal that vocal fold cancer progression is linked to cell surface receptor heterogeneity, a loss of laminin-binding integrins in cell-cell junctions and a flocking mode of collective cell motility. Mimicking the physiological movement of healthy vocal fold tissue with stretching or vibrations decreases oncogenic β-catenin and Yes-associated protein (YAP) nuclear levels in vocal fold cancer. Multiplex immunohistochemistry of vocal fold cancer tumours shows a correlation between the extracellular matrix composition, nuclear YAP and patient survival, concordant with vocal fold cancer sensitivity to oncogenic YAP-TEAD Hippo pathway inhibitors both in vitro and in vivo. Overall, our findings suggest that vocal fold cancer is a mechanically sensitive malignancy, and that the restoration of tumour mechanophenotype or YAP/TAZ targeting represents a tractable anti-oncogenic therapeutic avenue for vocal fold cancer.
    DOI:  https://doi.org/10.1038/s41563-025-02473-7
  17. Cell Syst. 2026 Feb 18. pii: S2405-4712(25)00340-0. [Epub ahead of print]17(2): 101507
      Are there general, systems-level principles guiding the evolution and design of natural or artificial sensory and signaling networks? Here, we argue that the signal transduction networks in living cells display important similarities in their organization and dynamical responses to both synaptic networks of brain cells and recent architectures of artificial neural networks. We propose that the key property of all of these networks-organization into multiple layers with hierarchically distributed timescales-is not accidental but rather reflects optimal processing of complex signaling and sensory inputs. We term this the hierarchical timescale hypothesis. We propose that the convergent evolution toward multi-step processing with "decreasing bandwidth" can also explain multiple properties of signaling networks, such as how a single input can control diverse outputs on different timescales and how noise and delay accumulation can be gracefully handled by the network.
    Keywords:  artificial neural network; hierarchical organization; multi-step processing; neuronal sensory network; signal transduction network; signaling network; synaptic network; systems-level principles; timescales
    DOI:  https://doi.org/10.1016/j.cels.2025.101507