bims-instec Biomed News
on Intestinal stem cells and chemoresistance in colon cancer and intestinal regeneration
Issue of 2026–07–12
eighteen papers selected by
Maria-Virginia Giolito, Université Catholique de Louvain



  1. Sci Adv. 2026 Jul 10. 12(28): eaeb8564
      Plasticity, the capability of tumor cells to go through phenotypic transitions, promotes colorectal cancer (CRC) progression and treatment resistance. Although plasticity is evident in advanced CRCs, little is known about plasticity in early-stage tumors and tumor stem cells. Here, we demonstrate that a plastic cell state (PCS) is present already in polyps from patients with familial adenomatous polyposis and in mouse intestinal adenomas, in which PCS is associated with PROX1+ tumor stem cells. We furthermore analyzed progressive plasticity upon loss of the canonical wingless-related integration site (Wnt) effector Tcf7 or Lef1 in Apc mutant mice. Deletion of either gene led to emergence of new plastic tumor cell populations, failure of leucine-rich repeat-containing G protein-coupled receptor 5 (Lgr5) tumor stem cell differentiation into enterocyte-like cells, enhanced Myc pathway activation, and increased tumor cell proliferation and tumorigenesis. Together, we demonstrate that PCS is associated with early CRC development and identify multiple potentially druggable mechanisms activated during progressive tumor cell plasticity.
    DOI:  https://doi.org/10.1126/sciadv.aeb8564
  2. bioRxiv. 2026 Jul 04. pii: 2026.06.30.735621. [Epub ahead of print]
      Colorectal cancer remains a major cause of cancer mortality, and most microsatellite stable tumors derive little benefit from immune checkpoint blockade. Here, we identify a microbiome-dependent mechanism that converts immune-refractory colorectal cancer into a more immunologically responsive state. Using orthotopic mouse models spanning distinct genetic and immunologic contexts, we show that a Helicobacter-containing microbiome suppresses primary tumor growth and limits metastasis. This protective state is associated with increased intratumoral lymphocyte infiltration and stronger effector programs. Mechanistically, microbial exposure induces MHC class II expression in colon cancer cells to promote anti-tumor immunity. Tumor-intrinsic loss of CIITA abrogates microbial protection, whereas enforced CIITA expression is sufficient to increase intratumoral T cell accumulation, restrict progression and metastasis, and sensitize microsatellite-stable tumors to PD-1 and CTLA-4 blockade. In human microsatellite-stable patient-derived organoids, increased cancer-cell MHC-II enhanced interactions with autologous immune cells and increased tumor cell apoptosis. Together, these findings define a microbiome-cancer cell antigen presentation axis that restrains metastasis and overcomes immunotherapy resistance in colorectal cancer.
    DOI:  https://doi.org/10.64898/2026.06.30.735621
  3. Oncogene. 2026 Jul 07.
      Claudin-1 (CLDN1), a tight junction protein overexpressed and mis-localized in colorectal cancer (CRC), plays a critical role in tumor progression, stemness, and therapy resistance. Integrative analyses of bulk and single-cell transcriptomic datasets revealed that CLDN1 is enriched in stem-like CRC cells, increases during metastatic progression, and is associated with microsatellite stable disease. High CLDN1 expression correlates with epithelial-to-mesenchymal transition and activation of oncogenic pathways including AKT/mTOR, Myc, and NF-κB. Here, we evaluate the therapeutic efficacy of an investigational humanized monoclonal antibody (H3L3) directed against overexpressed non-junctional CLDN1 in preclinical models of CRC. In xenograft mouse models and patient-derived organoids (PDOs), CLDN1 mAb significantly reduced tumor growth in CLDN1-expressing tumors. Mechanistically, CLDN1 mAb disrupted CLDN1-mediated signaling, notably inhibiting the AKT/mTOR pathway. CRISPR-mediated CLDN1 knockout abolished H3L3 efficacy, confirming target specificity. Transcriptomic analysis of PDOs treated with H3L3 revealed broad suppression of stemness, oncogenic signaling, and hallmark cancer pathways. Single-cell analysis further demonstrated that targeting CLDN1 modulates cellular plasticity, driving stem-like tumor cells toward a more differentiated epithelial phenotype. Together, these findings establish anti-CLDN1 monoclonal antibodies as a promising therapeutic approach for CRC. The selective binding of CLDN1 mAb to tumor-associated exposed and overexpressed CLDN1, along with its capacity to inhibit key oncogenic pathways and reprogram cancer cell states including tumors with different mutations conferring resistance to standard of care, underscores its potential to improve treatment outcomes in patients with CLDN1-expressing CRC.
    DOI:  https://doi.org/10.1038/s41388-026-03880-z
  4. Cell Rep. 2026 Jul 09. pii: S2211-1247(26)00699-6. [Epub ahead of print]45(7): 117621
      Argininosuccinate synthetase 1 (ASS1) is a rate-limiting enzyme in arginine biosynthesis, and its stability is regulated by TRAF2-mediated ubiquitination. Here, we report OTUB2 as a major deubiquitinase to stabilize ASS1, resulting increased arginine biosynthesis in colorectal cancer (CRC) cells; OTUB2 expression is elevated in CRC tissue, and patients with high OTUB2 expression exhibit shorter overall survival. As such, ectopic expression of OTUB2 promotes growth of CRC cells and xenograted tumors and accelerates cell migration and lung metastasis. Moreover, arginine deprivation induces marked expression of OTUB2 in CRC cells; mechanistically, arginine deprivation can activate AMPK, which in turn phosphorylates DDIT3, resulting its disassociation from C/EBPα and subsequent translocation into the cytoplasm and leading to increased binding of C/EBPα to the proximal promoter region of OTUB2 gene. Together, these data uncover a signaling pathway constituted of AMPK- DDIT3-C/EBPα-OTUB2-ASS1 to sense arginine deficiency and stimulate a metabolic compensatory pathway to sustain arginine homeostasis in CRC cells.
    Keywords:  ADT; AMPK; ASS1; C/EBPα; CP: cancer; CP: molecular biology; DDIT3; OTUB2; arginine deprivation therapy; ubiquitination
    DOI:  https://doi.org/10.1016/j.celrep.2026.117621
  5. Cancer Res. 2026 Jul 08.
      Cancer-associated fibroblasts (CAFs) play a crucial role in the tumor microenvironment (TME) by influencing tumor progression, metastasis, and therapy resistance. Accumulating evidence suggests that CAFs undergo senescence, which can impact their effects on the TME. Here, we developed a machine learning-based prediction model, the Cellular Senescence Prediction Model (CSPM), to accurately identify senescent CAFs (sCAFs) based on single-cell RNA sequencing data. In colorectal cancer (CRC), the abundance of sCAFs strongly correlated with impaired chemotherapy responsiveness and poor prognosis. In preclinical models, including subcutaneous tumors, patient-derived organoids (PDOs), patient-derived organoid xenografts (PDOXs), and orthotopic tumors, sCAFs mediated chemoresistance through the senescence-associated secretory phenotype (SASP), with IL6 and CXCL12 being key contributors. Macrophage-derived IL1B triggered CAF senescence through the IL1B-IL1R1 interaction, promoting the accumulation of sCAFs in tumors. Spatial transcriptomics and multiplex immunohistochemistry revealed colocalization of IL1B+ macrophages and IL1R1+ sCAFs in the tumor stroma. Functional studies using fibroblast-specific Il1r1 knockout mice further confirmed that macrophage-derived IL1B induces CAF senescence via IL1R1, leading to SASP-driven chemotherapy resistance. These findings highlight the critical role of sCAFs in CRC chemoresistance and suggest that targeting the IL1B-IL1R1 axis may offer a promising strategy to enhance chemotherapy efficacy in CRC.
    DOI:  https://doi.org/10.1158/0008-5472.CAN-25-4870
  6. Cancer Cell. 2026 Jul 09. pii: S1535-6108(26)00296-5. [Epub ahead of print]
      Colorectal cancer (CRC) metastases frequently recur due to minimal residual disease (MRD) and persistent micrometastases after therapy. Here, we performed spatial multimodal profiling using spot-level and high-resolution spatial transcriptomics, multi-regional whole-genome sequencing following laser-capture microdissection, and high-plex protein imaging to map 49 tumors from 19 patients, encompassing paired primary CRC and matched liver (CLiM) and lung (CLuM) metastases. Phylogenetic reconstruction revealed that liver micrometastases (CLiMi) arose from early clonal divergences and harbored a stem-like, quiescent state consistent with metastatic dormancy. Spatially, we uncovered distinct stromal barriers: macrometastases were encapsulated by myofibroblasts, whereas micrometastases were surrounded by immunosuppressive niches characterized by T cell exhaustion and distinct ligand-receptor signaling networks. Notably, we identified a CLiMi-specific six-gene signature associated with MRD status, disease-free survival, and chemotherapy resistance across multiple independent cohorts. These findings elucidate the spatial evolutionary landscape of CRC metastases and provide tissue-based spatially validated biomarkers for surveillance and therapeutic targeting.
    DOI:  https://doi.org/10.1016/j.ccell.2026.06.009
  7. Oncogene. 2026 Jul 07.
      In the tumor microenvironment (TME), cancer-associated fibroblasts (CAFs)-the dominant stromal component-actively shape cancer progression through exosome-mediated communication. Here, we identify hsa_circ_0003892 (circLDLR), a CAF-derived circRNA, as a key factor associated with poor prognosis in colorectal cancer (CRC). During interactions between CAFs and CRC cells, circLDLR is packaged into exosomes and transferred to tumor cells, where it enhances proliferation and metastasis primarily by reducing susceptibility to ferroptosis. Mechanistically, circLDLR stabilizes Polypeptide N-Acetylgalactosaminyltransferase 14 (GALNT14) by protecting it from ZNRF2-mediated ubiquitination and degradation. This stabilization promotes the O-GalNAcylation of Solute Carrier Family 7 Member 11 (SLC7A11) at Ser26, facilitating its membrane localization and thereby suppressing ferroptosis in CRC cells. Additionally, we demonstrate that the RNA-binding protein EIF4A3 facilitates circLDLR biogenesis within CAFs. Taken together, our study reveals that CAF-derived circLDLR confers ferroptosis resistance and promotes CRC progression via the GALNT14/SLC7A11 axis. Consequently, disrupting exosomal circLDLR transfer between CAFs and CRC cells may offer a promising therapeutic strategy for CRC. Schematic diagram of the mechanism by which CAF-derived circLDLR promotes CRC progression through ferroptosis regulation. In the tumor microenvironment, CAFs highly express circLDLR, whose biogenesis is facilitated by EIF4A3-mediated splicing of LDLR pre-mRNA. circLDLR is then packaged into exosomes and transferred to CRC cells. Within CRC cells, circLDLR specifically binds to GALNT14 and inhibits its ubiquitination and degradation mediated by ZNRF2, thereby enhancing GALNT14 protein stability. Stabilized GALNT14 promotes O-GalNAcylation of SLC7A11 at Ser26, facilitating its membrane localization and increasing cystine uptake. The elevated cystine metabolism enhances GSH production, thereby suppressing ferroptosis and driving CRC cell proliferation and progression.
    DOI:  https://doi.org/10.1038/s41388-026-03890-x
  8. Immunol Cell Biol. 2026 Jul 06.
      Mast cells (MCs), a type of granulocytic immune cell, exert contrasting effects on tumorigenesis. The anti- or pro-tumorigenic activity of MCs depends on the cancer type, tumor microenvironment, and MC localization within the tumor. Consequently, their role remains controversial and poorly understood across multiple cancer types, including colorectal cancer (CRC). Most proposed mechanisms underlying MC activity in CRC have focused on MC secretion of biological factors. In this study, we demonstrated that MCs transfer extracellular vesicles containing mRNAs and proteins to CRC cells. This process occurs through a tightly regulated mechanism that requires direct cell-cell contact, calcium signaling, and integrin-mediated interactions. Such requirements resemble aspects of immunological synapses observed between lymphocytes and cancer cells. The novel mode of intercellular communication between MCs and cancer cells described here may help refine our understanding of MC functions in cancer biology.
    Keywords:  colorectal cancer; extracellular vesicles; mRNA; mast cells
    DOI:  https://doi.org/10.1111/imcb.70147
  9. Cell Death Discov. 2026 Jul 11.
      Ferroptosis is an iron-dependent type of regulated cell death driven by lipid peroxidation, in which polyunsaturated fatty acids (PUFAs) in membrane phospholipids serve as key substrates. Here, we identify PUFA biosynthetic capacity as a key determinant of ferroptosis sensitivity under arachidonic acid (AA)-limited conditions. Lipidomic and stable isotope-tracing analyses in human lung adenocarcinoma cell lines revealed that H1299 cells that harbor wild-type KEAP1 and are sensitive to ferroptosis have unexpectedly impaired PUFA biosynthesis and consequently low PUFA levels. Nevertheless, ferroptosis sensitivity in H1299 cells is maintained under normal culture conditions owing to an exogenous supply of AA. Culturing cells in the B27 supplement, which provides only essential fatty acids such as linoleic acid, in the presence of 1% dialyzed FBS reduced AA-containing phospholipids and rendered H1299 cells resistant to ferroptosis. In contrast, A549 cells with a KEAP1 mutation retained ferroptosis sensitivity under B27 conditions, enabled by their endogenous PUFA-synthesizing capacity. Strikingly, pharmacological inhibition of FADS2 or deletion of ELOVL5 in A549 cells phenocopied H1299 cells, conferring resistance under B27-AO conditions that was reversed by AA supplementation. These findings demonstrate that ferroptosis susceptibility is dictated by intrinsic PUFA biosynthetic capacity under AA-limited conditions resembling physiological lipid availability. Therefore, intrinsic PUFA biosynthetic capacity should be considered in ferroptosis-based cancer therapies.
    DOI:  https://doi.org/10.1038/s41420-026-03240-6
  10. bioRxiv. 2026 Jun 29. pii: 2026.06.24.734305. [Epub ahead of print]
      Animals must allocate limited energetic resources across competing defense programs in response to infection. Here, we show that the conserved nuclear hormone receptor NHR-68 integrates fatty acid metabolism with the neural control of molecular and behavioral immunity in Caenorhabditis elegans . Acting in parallel with NHR-10, NHR-68 controls genes involved in polyunsaturated fatty acid (PUFA) metabolism. Loss of NHR-68 disrupts linoleic acid (LA) homeostasis, impairing pathogen avoidance behavior. Supplementation with LA restores avoidance, and fat-3 inhibition, which elevates LA, enhances pathogen avoidance, whereas loss of LA synthesis by fat-2 inhibition diminishes this behavior, indicating that LA promotes behavioral immunity. We further show that NHR-68 acts in the intestine to regulate linoleic acid homeostasis, and that changes in intestinal lipid metabolism influence an AWC-dependent pathogen-avoidance circuit through intestine-to-neuron communication. NHR-68 suppresses activation of the PMK-1/p38 MAPK and DAF-16/FOXO pathways, which mediate molecular immune responses. These findings identify a gut-brain transcriptional circuit that connects intestinal lipid metabolism to neural and immune outputs, revealing a mechanism by which the metabolic state coordinates behavioral and molecular defenses to optimize host protection.
    DOI:  https://doi.org/10.64898/2026.06.24.734305
  11. FASEB J. 2026 Jul 15. 40(13): e72114
      The metastatic potential of colorectal cancer (CRC) is a pivotal determinant of patient prognosis. Serine/threonine protein kinase 25 (STK25) is critically involved in diverse biological processes, and the function of STK25 in tumorigenesis and metastasis remains debatable across distinct tumor types. Here we identified that low STK25 expression was associated with increased tumor metastasis and poor survival in CRC patients. Functional experiments revealed that STK25 knockdown promoted CRC cells' epithelial-mesenchymal transition (EMT) and metastasis in vitro and in vivo. Mechanistically, STK25 depletion promoted migration and EMT progression through the TGF-β signaling pathway, and the kinase activity of STK25 was required to inhibit TGF-β signaling activation. These findings establish STK25 as a promising therapeutic target for intervening in TGF-β/SMAD2-mediated metastasis in CRC.
    Keywords:  EMT; STK25; TGF‐β; colorectal cancer; metastasis
    DOI:  https://doi.org/10.1096/fj.202504109R
  12. Cell Death Dis. 2026 Jul 04.
      Colorectal cancer (CRC) develops through a series of progressive genetic mutations, with alterations in APC and TP53 being the most frequently observed in the early stages. Although the loss of APC is recognized as a significant initiating event, the epigenetic processes through which the simultaneous inactivation of APC and TP53 facilitates the onset of colorectal tumorigenesis remain poorly characterized. To address this gap, we developed a human colon organoid model using CRISPR-Cas9 to achieve a double knockout of APC and TP53. Through extensive multi-omics profiling, we characterized the epigenetic landscape distinctive of early-stage CRC. We identified KIT, a receptor tyrosine kinase, as a critical oncogenic driver that is significantly upregulated in ΔDKO (APC and TP53 double knockout) organoids, thereby activating the MAPK and Wnt signaling pathways to augment proliferation and tumorigenesis. Furthermore, AP-1 transcription factors (FOS/JUN) regulate KIT expression via chromatin remodeling. Functional analyses indicated that KIT is integral to sustaining the elevated proliferation rates observed in ΔDKO organoids. These findings reveal a novel AP-1/KIT signaling axis that is central to the early progression of CRC, thereby presenting a promising avenue for therapeutic intervention.
    DOI:  https://doi.org/10.1038/s41419-026-09056-7
  13. Res Sq. 2026 Jul 05. pii: rs.3.rs-10164107. [Epub ahead of print]
      Endocytosis regulates receptor trafficking and signaling, yet its role in colorectal cancer (CRC) remains unclear. We analyzed transcriptomic data from 15,025 CRC tumors to evaluate gene expression in clathrin-mediated endocytosis (CME) and the endosomal sorting complexes required for transport (ESCRT) pathway. Pathway-level signatures were compared across consensus molecular subtypes (CMS) and examined in relation to oncogenic signaling programs, while individual endocytosis-related genes were evaluated for associations with clinical outcomes in an independent randomized trial cohort. Expression of CME and ESCRT pathways varied by CMS, with the highest expression in CMS4 and the lowest in CMS3. Endocytosis signatures correlated with multiple oncogenic signaling pathways, including MAPK, TGF-β, WNT, PI3K, NOTCH, and angiogenesis, suggesting broad links between endocytic activity and tumor biology. In the randomized phase III CALGB/SWOG 80405 trial, low expression of AP2M1 -a core adaptor in the CME complex-was significantly associated with longer overall survival and progression-free survival in patients treated with anti-EGFR agents, but not in those receiving anti-VEGF agents. These findings suggest that endocytic trafficking may be associated with oncogenic signaling and differential therapeutic benefit in patients with metastatic CRC. The endocytosis pathway reflects biologically relevant heterogeneity in CRC and may help guide therapeutic stratification.
    DOI:  https://doi.org/10.21203/rs.3.rs-10164107/v1
  14. Cell Rep. 2026 Jul 08. pii: S2211-1247(26)00752-7. [Epub ahead of print]45(7): 117674
      Cutaneous squamous cell carcinoma is a major cause of cancer-related mortality. Although immune checkpoint blockade improves outcomes, many patients fail to respond or develop resistance. Cancer-associated fibroblasts (CAFs) promote tumor progression and therapy resistance, but the signals that drive fibroblast reprogramming remain incompletely defined. Here, we show that thyroid hormone signaling, mediated by type 2 deiodinase (D2), regulates CAF activation. Single-cell transcriptomics and spatial RNA profiling identify a D2-positive fibroblast subpopulation that overlaps with immunomodulatory and matrix CAF states. RNA-seq analysis indicates that D2 supports activated and metabolically competent CAF programs. In vivo fibroblast-specific D2 deletion reduces CAF activation and limits tumor expansion. Spatial metabolomics further links D2-positive regions to metabolic remodeling of the tumor microenvironment and collagen-rich matrix deposition associated with aggressive tumor behavior. These findings support a role for D2-mediated thyroid hormone signaling in fibroblast reprogramming and suggest a therapeutic opportunity in cutaneous squamous cell carcinoma.
    Keywords:  CP: cancer; DIO2; cancer-associated fibroblasts; metabolic reprogramming; thyroid hormone signaling; tumor microenvironment cSCC
    DOI:  https://doi.org/10.1016/j.celrep.2026.117674
  15. bioRxiv. 2026 Jul 04. pii: 2026.06.30.735606. [Epub ahead of print]
      Colorectal cancer (CRC) remains a leading cause of cancer mortality, with most cases refractory to immunotherapy. Distinguishing tumor-induced from steady-state mucosal T cell responses has been a critical barrier to understanding antitumor immunity in CRC. Using orthotopic transplantation of CRC organoids with and without metastatic potential, combined with temporal T cell fate-mapping, we show that non-metastatic tumors elicit early recruitment of CD8αβ⁺ and CD4⁺ T cells that acquired cytotoxic and Th1-like programs, whereas pro-metastatic tumors induce a naïve-like, hypoactivated state. Tumor-infiltrating CD4 + T cells underwent clonal expansion, including clones recognizing microbial and dietary antigens. T cells in physical contact with tumor cells, identified by uLIPSTIC, were enriched for expanded and cytotoxic clones. Fate-mapped T cells from non-metastatic tumors suppressed tumor growth in an IFN-γ-dependent manner, whereas pro-metastatic tumor-derived T cells failed to do so. Mechanistically, pro-metastatic tumors downregulated MHCII, and Ciita targeting in non-metastatic organoids reduced CD4⁺ clonal expansion and led to tumor progression. Together, these findings define divergent early T cell trajectories associated with CRC metastatic potential, indicating that ineffective local immune engagement precedes metastatic dissemination.
    DOI:  https://doi.org/10.64898/2026.06.30.735606
  16. MethodsX. 2026 Dec;17 104016
      Mouse intestinal organoids are essential 3D models that retain host genetic characteristics and complex architecture, making them invaluable for intestinal biology research. However, traditional cultivation and histological processing often suffer from high operational complexity and sample loss during sectioning. This protocol provides an optimized workflow for the establishment of mouse intestinal organoid cultures and their subsequent preparation for immunohistochemistry and fluorescence staining. By refining the handling steps, the method significantly reduces total operational time while enhancing experimental quality. Furthermore, the protocol addresses common challenges in structural integrity, ensuring that the delicate crypt-villus morphology remains intact during the sectioning process. This comprehensive approach offers a robust framework for high-quality histological analysis, supported by a comparative evaluation with traditional methods to guide researchers in different experimental contexts.•Introduces a direct pellet-OCT embedding strategy that minimizes organoid transfer and sample loss during histological processing.•Enables high-quality frozen sections compatible with H&E, Alcian Blue, immunohistochemistry, and immunofluorescence staining.•Provides a practical alternative to conventional agarose-paraffin workflows while preserving organoid morphology and reducing processing complexity.
    Keywords:  Crypt; Embedding and sectioning; Organoids
    DOI:  https://doi.org/10.1016/j.mex.2026.104016
  17. Nat Genet. 2026 Jul 10.
      Phenotypically healthy cells frequently harbor somatic variants at cancer-associated genes, indicating that malignant transformation requires the selection of several alterations. Predicting which combinations of mutations, or co-mutations, exhibit oncogenic capacity requires identifying co-mutations that occur more or less frequently than expected. However, statistical frameworks to solve this problem are hampered by tumor heterogeneity and data availability. Here we curated putative oncogenic mutations in >70,000 human tumors from 119 subtypes, and designed a strategy to search for co-mutations based on in silico simulation of mutagenesis (SelectSim). Using this dataset and tool, we discovered and validated co-mutations across independent human cohorts, compared co-mutations across different tumor types and identified potential risk factors of metastatic progression. Notably, across several cohorts of phenotypically normal tissue samples, we show that, unlike individual oncogenic variants, significantly co-occurring mutations are largely cancer-specific and are observed rarely in healthy tissues, providing clues about the paths to tumorigenesis.
    DOI:  https://doi.org/10.1038/s41588-026-02661-4
  18. Nat Commun. 2026 Jul 04.
      Mucinous colorectal carcinoma (MUC CRC) metastasis to multiple organs, and to the peritoneum, is associated with poor prognosis. Disseminating MUC CRCs exhibit either conventional (apical-in) or inverted (apical-out) polarity that influence patient outcomes. Therefore, it is critical to identify how MUC CRC polarity is regulated. Here, we analyze patient-derived MUC CRC xenografts with either apical-in or apical-out polarity. Single-cell analyses reveal α2β1-integrin as a key collagen-binding receptor in these models. Collagen-α2β1-integrin interaction activates Src and upregulates SorLA, an endosomal sorting receptor. SorLA supports apical-in polarity by promoting integrin recycling and HER2/HER3 expression. We observe positive correlation between HER2, HER3 and SorLA in patient samples and higher HER2 expression in apical-in-presenting tissues. Clinically relevant HER2/HER3-targeting antibodies revert tumor sphere polarity, and impede collagen remodeling and adhesion to mouse peritoneum. This SorLA-integrin-HER2/HER3 axis could represent a MUC CRC-patient stratification approach and be relevant for other carcinomas with apical-out phenotypes.
    DOI:  https://doi.org/10.1038/s41467-026-75127-0