bims-enlima Biomed News
on Engineered living materials
Issue of 2026–07–12
forty-two papers selected by
Rahul Kumar, Tallinna Tehnikaülikool



  1. Adv Mater. 2026 Jul 09. e73986
      Designing polymers that combine tunable macromolecular architecture with complete chemical recyclability remains a fundamental challenge in sustainable polymer chemistry. Here, we report a δ-lactone copolymer platform synthesized via controlled ring-opening polymerization of δ-dodecalactone (δ7) and δ-valerolactone (δ0), enabling regulation of molecular architecture, mechanical response, and depolymerization pathways within a single polyester chemistry suitable for additive manufacturing (AM; or 3D printing). Segmental programming of δ7 and δ0 domains yields a synergistic combination of chain mobility and reversible crystalline reinforcement, producing recyclable, high-stiffness 3D-printing precursors with elastic modulus elevated by 2-3 orders of magnitude relative to prior δ-lactone materials. End-group functionalization generates methacrylate-terminated copolymers and acrylate-terminated macromonomers that undergo efficient photopolymerization without disrupting the depolymerizable backbone. Blending these components produces photocurable formulations with rheology governed by intrinsic polymer design, enabling compatibility with direct ink writing and digital light processing via temperature-mediated control of segmental crystallinity. This strategy eliminates external monomers or permanent crosslinkers, preserving chemical integrity and recyclability. Increasing molecular weight shifts the system into an entanglement-dominated thermoplastic regime for fused filament fabrication. Sequential thermal-catalytic unzipping exploits ceiling-temperature differences to recover δ7 and δ0 monomers with ∼95% efficiency even from crosslinked networks, with repolymerization yielding materials indistinguishable from virgin polymers.
    Keywords:  additive manufacturing; depolymerization; recycling; ring‐opening polymerization; δ‐lactone copolymers
    DOI:  https://doi.org/10.1002/adma.73986
  2. Nat Chem. 2026 Jul 08.
      The catalytic synthesis of many abiotic organic compounds is beyond the native capabilities of mammalian cells. Broadening the range of physiologically compatible, bond-forming catalytic reactions could enable the in situ synthesis of diverse molecules within living systems. By controlling these reactions in both specific cells and in physical space, this approach could offer transformative ways to modulate cellular behaviour and treat cells as distinct reaction vessels within organisms. Here we use genetic targeting to incorporate organic photocatalytic dyes onto and within cells to enable cell-type-specific and spatially defined catalysis. Photocatalysis can be restricted to individual cells using mild, green-to-red visible stimulation. The dyes catalyse cross-dehydrogenative C-H functionalization for the light- and cell-specific construction of C-C bonds in situ. Nuclear-localized assembly directs protein translocation with reversed transport directionality, illustrating the utility of this approach to both generate and enhance the function of chemical biology tools. We anticipate that the integration of synthetic catalysts with cell-specific targeting may lay an important foundation for future precision-directed abiotic chemical synthesis in living organisms.
    DOI:  https://doi.org/10.1038/s41557-026-02195-6
  3. Nature. 2026 Jul;655(8122): 372-380
      Hydrogels are widely used in biomedical interfaces, in which effective gas exchange (for example, O2, CO2) within a water-rich environment is essential. However, hydrogels show intrinsically limited air exchange efficiency, owing to the low solubility (C) and diffusivity (D) of non-polar gases in the polar water medium1. This limitation poses a substantial bottleneck in long-term applications, such as wearable health monitors2-7 and tissue engineering8-12. Existing methods13-16 to enhance air permeability suffer from poor robustness and/or an inherent trade-off between permeability and water content (for example, <50 vol%). Here we introduce a viscoelastic phase separation17 (VPS)-enabled strategy to create a non-collapsible, air-rich network in high-water-content hydrogels, achieving a record-high oxygen permeability of 185 barrer with 70 vol% water-a tenfold increase compared with pristine hydrogels. VPS, a ubiquitous phenomenon in soft matter, is used to drive hydrophobic, dry gas particles within a hydrophilic, wet medium into a thin, stable three-dimensional network. This approach allows the facile and scalable fabrication of air-permeable hydrogels across diverse chemistries and form factors. Physiological tests over a 10-day continuous wear condition confirmed their effectiveness in preventing fluid accumulation and maintaining skin health. This strategy paves the way for hydrogels in long-term biomedical applications in which efficient and sustained air exchange becomes critical.
    DOI:  https://doi.org/10.1038/s41586-026-10712-3
  4. Curr Opin Microbiol. 2026 Jul 10. pii: S1369-5274(26)00083-4. [Epub ahead of print]93 102789
      The field of engineered living materials (ELMs) aims to create self-regenerative, self-assembled, and multifunctional materials that mimic natural biomaterials. Novel ELMs can be produced by engineering biomolecules that are naturally secreted and displayed on bacterial cell surfaces. Surface-layer (S-layer) proteins are a class of proteins that form a two-dimensional paracrystalline lattice on the surface of many prokaryotes. These proteins provide a secretion, surface-anchoring, and high-density display platform that can be exploited for material formation. In this review, we discuss two strategies to engineer S-layer proteins for ELMs by looking at their state of the art, analyzing their advantages and disadvantages, and discussing their challenges and opportunities.
    DOI:  https://doi.org/10.1016/j.mib.2026.102789
  5. Adv Sci (Weinh). 2026 Jul 10. e76493
      Size exclusion within biological hydrogels imposes a fundamental constraint on the design of nanocarriers, limiting the transport of cargo-loaded and structurally complex materials through mucus barriers. While surface passivation strategies are commonly used to improve compatibility, they do not address steric limitations imposed by the polymer network. Here, we introduce mechanical flexibility as an independent materials design parameter to expand the functional transport window of nanocarriers in mucus. Using programmable DNA origami to decouple flexibility from size and surface chemistry, we show that increased structural compliance enhances transport under steric confinement by facilitating passage through confined network pores. When surface-driven aggregation dominates, passivation is required to restore transport, after which flexibility provides additional gains. Together, these results establish mechanical flexibility as a general materials design strategy for improving transport under size-constrained conditions, with implications for nanocarrier engineering across biological barriers.
    Keywords:  DNA nanotechnology; oral drug delivery; single particle tracking; single‐particle tracking
    DOI:  https://doi.org/10.1002/advs.76493
  6. Sci Adv. 2026 Jul 10. 12(28): eaec3482
      Elastomers can be engineered as stretchable, compliant materials capable of bearing load, making them attractive for soft robotics, wearable electronics, and biomedical devices. Unfortunately, single-network elastomers are constrained by a stiffness-toughness compromise. While multinetwork elastomers can mitigate this limitation, tough multinetwork elastomers typically exhibit a limited fatigue resistance. Here, we demonstrate that double-network granular elastomers (DNGEs) composed of stiff elastomer microparticles connected through a softer second network combine toughness and fatigue resistance. Their locally varying composition and structure enable DNGEs to efficiently deconcentrate stress and repetitively dissipate energy if stretched to moderate strains, imparting them a good fatigue resistance. Leveraging the three-dimensional printability of DNGEs, we spatially vary the composition to combine stiffness, toughness, and fatigue resistance within one elastomer. We anticipate that DNGEs will unlock opportunities in soft robotics, wearables, and biomedicine, where current soft materials remain limited by the stiffness-toughness-fatigue resistance trade-off.
    DOI:  https://doi.org/10.1126/sciadv.aec3482
  7. ACS Nano. 2026 Jul 10.
      Cells use a combination of membrane-bound and membrane-less compartments to dynamically orchestrate internal biochemical processes and sustain intracellular communication. Recapitulating the hierarchical integration and interplay between these physically and chemically diverse structures is required to enhance the functionalities of synthetic cells and other advanced biomimetic systems. Here, we describe the use of synthetic DNA condensates to selectively uptake and spatially organize lipid vesicles, interacting with the condensates thanks to cholesterol-DNA anchors. By modulating anchor density, the liposomes can be programmably localized on the surface or interior of the condensates, while base-pairing selectivity can be leveraged to target individual internal domains in multiphasic condensates. The embedded liposomes can be released by adding a nucleic acid trigger and captured by a second condensate population, thus imitating extracellular vesicles in their ability to support long-range cellular communication. This modular platform demonstrates the potential of DNA-based condensates to program the spatial distribution of membranous subcompartments and to support dynamic cargo-handling capabilities. These features are valuable for engineering cell mimics, microreactors, and delivery systems.
    Keywords:  Compartmentalization; Condensates; DNA nanotechnology; LLPS; Liposomes
    DOI:  https://doi.org/10.1021/acsnano.5c16730
  8. Bioinformatics. 2026 Jul 01. pii: btag219. [Epub ahead of print]42(Supplement_1):
       MOTIVATION: Recent dynamic lineage tracing technologies use genome editing to induce heritable mutations, or edits, that accumulate across successive cell divisions. These edits are measured using single-cell sequencing or imaging, providing data to reconstruct cell lineages at single-cell resolution. Current computational approaches to infer cell lineage trees, or phylogenies, from these data perform two separate steps: (i) Identify each cell's edits (genotype) from the raw sequencing or imaging data; (ii) Infer a cell lineage tree from the cell genotypes. However, genotyping cells is an inexact process and genotype errors can yield an inaccurate lineage tree. For example, using fluorescence based-imaging to measure edits results in a high fraction (≈25%-50%) of uncertain or erroneous genotypes.
    RESULTS: We introduce Lineage Analysis via Maximum Likelihood with PRobabilistic Observations (LAML-Pro), an algorithm that jointly infers cell genotypes and a cell lineage tree. LAML-Pro is based on the Probabilistic Mixed-type Missing Observation (PMMO) model, which we derive to describe both the genome editing and genotype observation processes. LAML-Pro constructs lineage trees from thousands of cells in under an hour by leveraging the sparsity of transitions under the PMMO model. On simulated data, we demonstrate that LAML-Pro corrects genotype errors and infers substantially more accurate trees than existing methods which are vulnerable to genotype errors. Applied to data from two recent imaging-based lineage tracing systems, LAML-Pro reduces genotype errors by 5-fold and produces more spatially coherent lineage trees compared to existing methods.
    AVAILABILITY AND IMPLEMENTATION: LAML-Pro is implemented in C++ and is available as both a command-line interface and as a Python library at: github.com/raphael-group/LAML-Pro.
    DOI:  https://doi.org/10.1093/bioinformatics/btag219
  9. Nat Mater. 2026 Jul 10.
      Hydrogels are widely used in biomedical and bioelectronic applications owing to their tissue-like properties, including biocompatibility, softness and three-dimensional architecture. In recent years, semiconducting behaviour was demonstrated in hydrogels through the network design of π-conjugated polymers, extending their potential to advanced electronic applications such as transistors. In this Review, we provide an overview of the design, fabrication, characterization and benchmarking standards of π-conjugated hydrogel semiconductors, a rapidly evolving field requiring interdisciplinary knowledge across organic electronics, electrochemistry and soft materials. We demonstrate how to understand and regulate ion and electron transport, as well as their interactions, both thermodynamically and kinetically, in these π-conjugated supramolecular systems. Finally, we envision the potential of these materials to advance spatiotemporal biological research, wearable healthcare, implantable medicine and beyond.
    DOI:  https://doi.org/10.1038/s41563-026-02638-y
  10. Cell Biomater. 2026 Jun 16. pii: 100404. [Epub ahead of print]2(6):
      Almost every cell in vivo is surrounded by the extracellular matrix (ECM), which contributes to cell and tissue fate. Biomaterials, especially engineered hydrogels, have emerged as platforms to recreate the biochemical and mechanical properties of ECM. Yet most approaches still assume that cells directly respond to the engineered hydrogel, although there is increasing evidence that numerous cell types rapidly deposit newly synthesized (nascent) ECM (nECM). Studies have now shown that this nECM accumulates at the cell-hydrogel interface, where it also contributes to a cell's fate. In this perspective, we first highlight key studies describing the nECM as a regulator of cell fate. Next, we provide guidelines based on physicochemical principles for studying nECM through spatiotemporal mapping, mechanical/structural measurements, biochemical characterization, and functional assays. Finally, building upon existing hydrogel engineering tools, we propose chemical- and protein-engineering approaches to specifically engineer the nECM to more precisely control cell-matrix interactions.
    Keywords:  Extracellular matrix; cell culture; cell-matrix interactions; hydrogel; protein engineering
    DOI:  https://doi.org/10.1016/j.celbio.2026.100404
  11. Mater Today Bio. 2026 Aug;39 103368
      Light-assisted 3D-bioprinted hydrogels are at the forefront of tissue engineering and biomedical manufacturing due to their precise spatiotemporal controllability and tunable physicochemical properties. However, the mechanical requirements of target organ tissue pose challenges to 3D-bioprinted bio-functional tissue, which must balances biocompatibility, printability, and mechanical strength to replicate the native regenerative microenvironments for functional restoration. This review explores photosensitive polymers with bond chemistry for strengthening hydrogel-based tissue constructs, offering an examination of their potential for facilitating tissue reconstruction through light-assisted 3D bioprinting techniques. We compare the cross-linking chemistry, bond energetics, and resulting mechanics of natural and synthetic hydrogels. Additionally, various light-assisted 3D bioprinting methods for tough hydrogels and engineered living systems are summarized and compared in terms of their practical applications. Importantly, we highlight the critical challenge of enhancing mechanical toughness while balancing the printability, toughness, and biocompatibility of hydrogels via the use of covalent bonds, dynamic covalent bonds, reversible non-covalent interactions, and hybrid bond networks. Furthermore, we discuss emerging applications of light-printed tough hydrogel scaffolds in regenerative medicine and in cartilage, bone, tendon, skull, musculoskeletal, and dental applications. Future prospects and challenges associated with methods of toughing 3D-bioprinted hydrogels are also discussed for guiding future biomedical engineering efforts.
    Keywords:  3D bioprinting; Biomedical applications; Photopolymerized hydrogels; Tough hydrogels
    DOI:  https://doi.org/10.1016/j.mtbio.2026.103368
  12. Chembiochem. 2026 Jul 14. 27(13): e70450
      Membrane-associated biomolecules, primarily proteins, are key enablers of communication, responsiveness, and complexity in natural living cells. Aiming to mimic these capabilities, there is growing interest in equipping bottom-up synthetic cells with membrane-associated biomolecular components. In this review, we focus on how proteins and nucleic acids have been associated with synthetic cell membranes, particularly lipid vesicles, to enable the transmission of signals across the membrane. We discuss strategies for anchoring these biomolecules into lipid bilayers and review how they can enable essential signalling mechanisms in synthetic cells, including cell tethering, the generation and fusion of vesicles, and signal transmission and transduction. We highlight how proteins offer native biological functionality, while nucleic acids may bring more modularity and control. Advancing this area will be essential for realising synthetic systems capable of studying natural communication mechanisms and unlocking applications in biosensing, therapeutics, and synthetic tissue engineering.
    Keywords:  artificial cell; biomolecule; biosensor; cell signalling; lipid bilayer; membrane; nucleic acid; synthetic biology; vesicle
    DOI:  https://doi.org/10.1002/cbic.70450
  13. Nat Chem Eng. 2026 Jun;3(6): 328-339
      Replicating the intricate hierarchical architecture of natural vascular networks, especially at capillary-scale resolution, remains a pivotal challenge in organ fabrication. Here we present a machine learning-enhanced hybrid bioprinting strategy that combines high-resolution aerosol jet printing of sacrificial materials and high-throughput extrusion printing of tissue matrices. This integrated approach enables sub-10 μm of resolution, achieving capillary-like channels and allowing on-demand modulation of vessel diameters in real time. Constrained Bayesian optimization rapidly identify optimal printing parameters, ensuring reliable, high-fidelity attainment of target channel sizes without exhaustive trial-and-error. This streamlined workflow supports the fabrication from 1D conduits to 3D multibranch hierarchical networks with tunable geometries. Endothelial cells seeded into these channels form continuous, functional monolayers, significantly reducing permeability while maintaining high cell viability and proliferation. By transcending the resolution limits of conventional sacrificial printing, this bioprinting method establishes a new route for producing biomimetic vasculature. Its unique combination of rapid optimization, real-time tunability, and microcapillary-scale precision holds exceptional promise for tissue engineering, regenerative medicine, and drug discovery.
    DOI:  https://doi.org/10.1038/s44286-026-00396-x
  14. Nucleus. 2026 Dec 31. 17(1): 2690847
      Mechanical cues, ranging from matrix mechanical properties to dynamic mechanical loading, can be transmitted via structural proteins and signaling molecules to the nucleus to reorganize nuclear architecture and modulate chromatin accessibility. This mechanical regulation plays an important role in tissue regeneration and disease development. To gain deeper insights into the mechanical regulation of chromatin organization, it is essential to develop technologies that can apply mechanical inputs and characterize the resulting changes in nuclear structure and chromatin organization. Here, we review multidisciplinary technologies and tools that enable mechanical perturbation of the nucleus and the characterization of nuclear and chromatin responses. We highlight how perturbations such as matrix topography, confinement, stiffness, viscoelasticity, and dynamic loading can be used to apply mechanical cues to cells. We also discuss how imaging-based techniques, sequencing platforms, and computational approaches can be integrated to characterize nuclear architecture and chromatin organization in response to these mechanical stimuli.
    Keywords:  Mechanobiology; biomaterials; epigenetic regulation; microfluidics; nuclear mechanics; single cell imaging
    DOI:  https://doi.org/10.1080/19491034.2026.2690847
  15. Mater Horiz. 2026 Jul 08.
      DNA hydrogels are widely explored in biomedical research for their programmability and soft tissue-mimicking mechanics. However, their application in load-bearing implants is restricted by insufficient mechanical robustness, especially for mimicking the biomechanics of the nucleus pulposus, which is a naturally occurring functional tissue essential for spinal flexibility and shock absorption in the intervertebral disk. Here, we report a novel single-pot, two-step fabrication strategy in which long DNA strands produced by rolling circle amplification form an initial viscoelastic network that is subsequently reinforced through controlled thermal self-assembly of proteins, physically stapling the DNA chains into a mechanically tunable hybrid matrix. This synergistic protein reinforcement enables precise control over the morphological and mechanical properties of DNA hydrogels while improving stability under enzymatic and pH stress. The reinforced hydrogels maintain structural integrity under complex deformation and sustained compression in ex vivo nucleus pulposus models and exhibit pressure-dependent drug release. Overall, this study establishes protein reinforcement within viscoelastic gel networks as a distinct materials design strategy for creating programmable biomaterials that integrate molecular precision with mechanical resilience for use in mechanically demanding biological environments.
    DOI:  https://doi.org/10.1039/d6mh00586a
  16. Nat Commun. 2026 Jul 08.
      Engineered small RNAs (sRNAs) enable programmable gene knockdowns and support metabolic engineering and multiplex regulation in model bacteria. Still, precise, tunable, and multiplex gene repression remains a challenge in synthetic biology. Common tools can impose genetic burden, depend on host RNA factors, or do not transfer well across species. Here we present MORTISE (Multiplex, ORthogonal Translation Interference SystEm), a compact Cas6f-based platform for programmable translational repression in Gram-negative bacteria. The system functions without host Hfq or RNases and operates robustly in Escherichia coli and Pseudomonas putida. We demonstrate repression in both species using chromosomal reporter assays, with performance improving when guide and target transcription are matched and when the translation initiation region is targeted. Single-promoter multiplexing enables simultaneous knockdowns and a cloning toolbox facilitates assembly of up to nine guides in a single step. Finally, MORTISE is leveraged to boost malonyl-coenzyme A-dependent production in P. putida, supporting pathway balancing.
    DOI:  https://doi.org/10.1038/s41467-026-75135-0
  17. bioRxiv. 2026 Jun 29. pii: 2026.06.26.734876. [Epub ahead of print]
      Precisely modifying proteins at multiple sites in their native, folded structures offers unique opportunities to answer molecular and cellular-level biological questions. Here, we present a genetic code expansion strategy for site-specific integration of a fluorophore-quencher pair comprising two non-canonical amino acids-acridonylalanine (Acd) and methyltetrazinyl phenylalanine (Tet) - into a protein expressed in E. coli . The Acd and Tet pair requires no post-translational labeling, and quenching can be switched off by biorthogonal or photochemical reactions of Tet for convenient internal control experiments. Mechanistic studies based on Stern-Volmer quenching, fluorescence lifetime measurements, and "proline ruler" peptides established the distance dependence of quenching. As proof-of-concept, we applied this strategy to study: 1) calmodulin, a calcium-sensing protein, 2) RecA, a DNA damage sensor in bacteria, and 3) LexA, a transcriptional repressor whose activation by RecA governs acquired antibiotic resistance in bacteria. Using these proteins, we demonstrate that dual Acd/Tet labeling provides molecular-level insights into protein dynamics, enables high-throughput drug screening, and advances tools for studying protein structure-function relationships.
    DOI:  https://doi.org/10.64898/2026.06.26.734876
  18. Proc Natl Acad Sci U S A. 2026 Jul 14. 123(28): e2532667123
      Engineering organized microvascular networks remains a critical challenge in tissue engineering and regenerative medicine. While biochemical approaches for patterning angiogenesis via growth factor delivery have shown promise, their inability to pattern sustained growth factors with spatiotemporal control limits effectiveness. Here, we demonstrate that dynamically patterned mechanical forces enable precise spatiotemporal control over angiogenic sprouting. We developed a magnetically actuated human vessel-on-a-chip platform that integrates a perfusable endothelialized microchannel within a collagen matrix and allows noninvasive and tunable mechanical stimulation across three spatial dimensions and time (4D). Using an automated 3-axis actuator, we systematically investigated how strain magnitude, frequency, and direction modulate endothelial cell behavior and vessel morphogenesis. Dynamic mechanical stimulation at physiological strain magnitudes (5 to 15%) enhanced endothelial alignment and barrier function while promoting angiogenesis in a strain magnitude-dependent manner: lower dynamic strain (5%) maximized sprout initiation, whereas higher dynamic strain (15%) promoted elongation of sprouts. Sequential reorientation of strain direction reprogrammed sprouting trajectories along X, Y, and Z directions, generating complex sprout geometries such as L-shaped branches. RNA sequencing revealed mechanically induced transcriptional profiles distinct from unstimulated controls, characterized by upregulation of genes associated with angiogenesis, mechanotransduction, and extracellular matrix remodeling. Functional perturbation of PIEZO1 reduced strain-induced sprouting without altering barrier function, indicating that dynamic mechanical stimulation engages multiple mechanotransduction pathways to regulate angiogenesis. Collectively, these findings establish a strategy for spatiotemporally controlled angiogenesis through 4D force patterning to program vascular morphogenesis while preserving function. This approach provides a foundation for engineering hierarchically organized vascular networks for tissue regeneration.
    Keywords:  angiogenesis; biofabrication; mechanobiology; tissue engineering; vasculature
    DOI:  https://doi.org/10.1073/pnas.2532667123
  19. bioRxiv. 2026 Jul 03. pii: 2026.07.02.736090. [Epub ahead of print]
      Generating bioengineered cartilage that recapitulates the depth-dependent phenotype, structure, and function of native articular cartilage remains a challenge. While cartilage is rich in aggrecan and type II collagen, proper function depends on depth-dependent protein expression. Superficial zone chondrocytes (SZCs) secrete proteoglycan-4 (PRG4) to lubricate the cartilage surface. Deep zone chondrocytes produce type X collagen (COLX) to support compressive loading and load transfer to subchondral bone. We previously demonstrated that passaged full-thickness chondrocytes (FTCs) and zonal chondrocytes can re-express cartilage and zone-specific markers following scaffold-free three-dimensional (3D) culture in redifferentiation media. However, in the absence of an instructive matrix, cells expressed low levels of zone-specific proteins and exhibited limited depth-dependent organization. We hypothesize that synthetic extracellular matrix with zone-specific microenvironmental cues will guide zonal differentiation . To this end, passaged primary bovine chondrocytes were encapsulated in a soft, hyaluronan (HA)-based, cell-adhesive, and protease-degradable hydrogel established via bioorthogonal tetrazine (Tz) ligation with norbornene (Nb). When supplemented with TGFβ3, FTCs deposited aggrecan and type II collagen with minimal type I collagen. Application of interfacial tetrazine ligation with trans -cyclooctene (TCO) during cell culture resulted in matrix stiffening, leading to upregulation of COLX expression. Conversely, SZCs cultured in soft hydrogels exhibited the greatest PRG4 expression. Establishment of a trilayered construct with region-specific stiffness via the diffusion-controlled reaction promoted PRG4 and COLX expression in defined zones. Together, these findings demonstrate that tunable HA-based hydrogels can enhance zone-specific chondrocyte phenotypes and promote the formation of zonally organized cartilage.
    DOI:  https://doi.org/10.64898/2026.07.02.736090
  20. Sci Adv. 2026 Jul 10. 12(28): eaee9688
      Developing a reliable, long-term electrical interface for implanted bioelectronics is essential for chronic diagnostics, therapeutics, and device maintenance. Here, we introduce the implantable bioelectronic outlet (IBO), a soft, tissue-like electronic interface that enables on-demand, direct ohmic connection between implanted electronics and external devices. IBO is composed of a conducting polymer-coated low-density polymer matrix that is jacketed with a hydrophobic elastomer, allowing repetitive insertion of external electrical contacts without crack propagation. IBO was functional after 1 year of implantation with minimal tissue effects. We demonstrate that IBO enables high-fidelity, bidirectional transmission of signals and power, including low-voltage neurophysiological signals, high-speed digital signals, neurostimulation protocols, and efficient high-current power delivery, validated in small and large animal model studies. The IBO provides a robust and scalable platform for safe, direct, and durable electrical interfacing with implantable bioelectronics.
    DOI:  https://doi.org/10.1126/sciadv.aee9688
  21. ACS Appl Mater Interfaces. 2026 Jul 10.
      Conductive gels are promising materials for on-skin wearable bioelectronics owing to their softness, ionic conductivity, and tissue-like interfacial characteristics. However, conventional preformed conductive gels often cannot fully accommodate complex and dynamically deforming skin surfaces, thereby limiting their interfacial adhesion and signal-transmission performance. Inspired by the reversible mechanical switching behavior of sea cucumbers, we developed a thermoresponsive ionic biogel (IBG) composed of gelatin, water, and 1-ethyl-3-methylimidazolium ethyl sulfate. In this system, gelatin forms a physical network with reversible sol-gel transition behavior, while the ionic liquid modulates the network through hydrogen bonding and electrostatic interactions and provides mobile ions for charge transport. This design integrates mechanical compliance, skin adhesion, water retention, and electrical functionality. The IBG can be directly coated onto skin in a flowable state and subsequently undergo mild in situ gelation, forming a soft, adaptive, and ion-conductive bioelectronic interface. The optimized IBG43 exhibits a low modulus, high stretchability, strong adhesion, and good environmental stability, together with stable strain sensing, reliable electrophysiological signal acquisition, and potential for self-powered sensing. This work presents a simple and effective strategy for constructing in situ formable skin-interfaced bioelectronic materials.
    Keywords:  flexible bioelectronics; gelatin; in situ gelation; ionic biogel; ionic liquid; skin-conformal biointerface
    DOI:  https://doi.org/10.1021/acsami.6c06889
  22. bioRxiv. 2026 Jun 29. pii: 2026.06.26.734902. [Epub ahead of print]
      The immune system uses paracrine signaling to spatially confine potent responses such as inflammation. A bio-orthogonal synthetic paracrine system could enable engineering of analogous multicellular circuits in which different cell types coordinate their functions in a spatially organized fashion. Here, using the plant hormone auxin as a bio-orthogonal chemical signal, we introduce programmable paracrine circuits that distribute sensing and effector functions to different cell types to spatially restrict responses in mouse xenografts. Cells engineered to express auxin biosynthetic genes generated auxin-dense regions with tunable length scales in vivo. This localized signaling ability enabled design of a multicellular sentinel-effector system, in which THP-1 sentinel cells conditionally produce auxin in regions expressing the tumor-specific antigen EGFRvIII, and Jurkat effector cells respond by locally modulating the activity of a chimeric antigen receptor (CAR). This two-cell type system was able to achieve localized activation of engineered effector cells in vivo. These results establish a foundation for engineering multicellular therapeutic systems that focus responses in specific tissue contexts or disease sites.
    DOI:  https://doi.org/10.64898/2026.06.26.734902
  23. Nat Commun. 2026 Jul 04.
      Understanding how living organisms spontaneously develop complex functional structures motivates new strategies in engineering design. Here, we introduce a decentralized generative model based on morphogenesis to autonomously grow mechanical structures with controlled tensorial properties. By adapting Turing's reaction-diffusion concept through anisotropic diffusion, our approach enables the local emergence of microstructures exhibiting tailored stiffness and anisotropy, achieving target orthotropic tensors without adjoint or topology optimization loops. The synthesis of these structures relies on a database linking morphogenetic parameters to effective elastic tensors obtained through homogenization techniques. We experimentally demonstrate this concept through a mechanical cloaking example, validating our method's capability to independently control local anisotropy and rigidity, and effectively conceal structural defects from mechanical fields. This approach circumvents the iterative global solves required by topology optimization, while preserving local control over anisotropy and stiffness across large design domains.
    DOI:  https://doi.org/10.1038/s41467-026-74173-y
  24. Proc Natl Acad Sci U S A. 2026 Jul 14. 123(28): e2525718123
      mRNA-based cancer vaccines offer a modular and safe platform to elicit antitumor immunity, yet their efficacy is often limited by inefficient mRNA delivery and inadequate dendritic cell (DC) activation, both of which are essential for initiating robust cytotoxic T cell responses. Inadequate innate immune activation coupled with poor antigen presentation further diminishes their effectiveness, particularly in immunologically "cold" tumors. While stimulator of interferon genes (STING) agonists can enhance DC maturation and cross-presentation, their therapeutic utility is constrained by poor intracellular delivery and limited colocalization with tumor antigens. In this study, we developed a lipid nanoparticle (LNP) platform via high-throughput screening of ionizable lipids for potent mRNA delivery to DCs both in vitro and in vivo. To amplify immune activation, we coencapsulated the STING agonists c-di-AMP (AMP) and manganese (Mn2+) together with tumor antigen-encoding mRNA into the lead LNP formulation. This codelivery strategy synergistically activated type I interferon signaling, upregulated costimulatory molecules, enhanced antigen presentation, and elicited potent tumor-specific T cell responses and superior antitumor efficacy. Our results demonstrate that integrating innate immune stimulation with mRNA-LNP delivery provides a promising strategy to overcome current limitations in mRNA vaccine efficacy and to improve cancer immunotherapy outcomes.
    Keywords:  cancer; immunotherapy; mRNA; nanoparticle
    DOI:  https://doi.org/10.1073/pnas.2525718123
  25. bioRxiv. 2026 Jul 02. pii: 2026.07.01.735572. [Epub ahead of print]
      Cell penetrating thyclotides (CPTs) are synthetic molecules that promote highly efficient cellular uptake and endosomal escape of bioactive peptides. While peptides are valuable as medicinal agents, their translation to therapies is often limited by their inability to cross cell membranes. CPTs have a unique combination of chiral tetrahydrofurans and polar sidechains within a molecular scaffold that can be optimized to efficiently deliver peptide cargo into cells. The cellular uptake and endosomal escape of two peptides with anticancer biological activities but low bioavailabilities were remarkably improved after conjugation to a CPT. Using CPTs to overcome barriers to cellular uptake represents a new direction for the intracellular delivery of bioactive molecules, and will accelerate drug development for new medical therapies.
    DOI:  https://doi.org/10.64898/2026.07.01.735572
  26. bioRxiv. 2026 Jul 01. pii: 2026.06.29.735318. [Epub ahead of print]
      Implant-associated infections are driven by bacterial biofilm formation and remain difficult to eradicate using conventional antibiotic-based strategies. Here, we present a dynamically actuated reconfigurable topographical surface (DARTS) that integrates intrinsically bactericidal nanoscale surface topography with programmable mechanical actuation to achieve durable, antibiotic-free infection control. Using a scalable bottom-up nanofabrication strategy, we generate tunable wrinkled MXene topographies that exhibit contact-mediated bactericidal activity against both Gram-positive and Gram-negative bacteria without chemical leaching. Integration with a soft robotic actuator enables reversible modulation of surface geometry, which synergistically enhances bacterial removal and killing, resulting in near-complete disruption of mature biofilms. Dynamic actuation further sensitizes released bacteria to antibiotic treatment. In a mouse subcutaneous implant infection model, DARTS with actuation achieves sustained suppression of bacterial burden and markedly improves host tissue outcomes. Remote, noninvasive actuation using near-infrared laser stimulation further highlights the translational potential of this platform for implantable antibacterial applications.
    Significance Statement: Implant infections are difficult to treat because bacteria form biofilms that protect them from antibiotics and the immune system. Current materials often rely on chemical release, which can lose effectiveness over time. Here, we present a new surface that both kills bacteria and removes them. The surface uses nanoscale features to physically damage bacterial cells, while dynamic motion clears attached bacteria and biofilms. This allows continuous, chemical-free control of infection. In a mouse implant model, the system greatly reduced bacterial burden and improved tissue healing. This work introduces a new way to control bacteria using dynamic surface design and could be applied not only to medical implants but also to environmental, food, textile, and marine systems.
    DOI:  https://doi.org/10.64898/2026.06.29.735318
  27. Trends Biotechnol. 2026 Jul 06. pii: S0167-7799(26)00256-8. [Epub ahead of print]
      Biomaterials are evolving from passive scaffolds to responsive platforms, yet most lack true feedback regulation. A shift toward adaptive biomaterial systems that integrate sensing, computational processing, and dynamic actuation directly within a macromolecular network could transform these multifunctional biomaterials into precision platforms capable of regulating biological processes in real time.
    Keywords:  biointerfaces; biomaterials; closed-loop systems; feedback control; precision therapy
    DOI:  https://doi.org/10.1016/j.tibtech.2026.06.016
  28. Nat Microbiol. 2026 Jul 07.
      Bacteria residing in biofilms are embedded in an extracellular matrix. Whereas biofilm formation is well studied, less is known about biofilm dispersion, although enzymatic extracellular matrix degradation is suspected to play a key role. Here we show that Bacillus subtilis biofilms can alternatively eject a specific cell type, locally and anisotropically, using mechanical forces arising from a self-generated hydrogel. Single-cell resolution imaging combined with mathematical modelling, and chemical and genetic perturbations, show that the production of the extracellular poly-γ-glutamic acid (γ-PGA) polymer is necessary to drive this cell ejection. Specifically, osmotic pressure from the γ-PGA hydrogel propels interior cells through the outer layers to break free from the biofilm. We demonstrate control over this process through γ-PGA modulation such that biofilm dispersion can be either inhibited or promoted. Forceful ejection driven by γ-PGA has so far only been described in marine organisms such as jellyfish. Our discovery of biofilm cell ejection via γ-PGA thus reveals not only a previously uncharacterized biofilm dispersion mechanism but also an unexpected mechanistic parallel to evolutionarily distant Cnidaria.
    DOI:  https://doi.org/10.1038/s41564-026-02413-4
  29. ACS Mater Au. 2026 Jul 08. 6(4): 716-728
      Hydrogels are everyday materials characterized by their remarkable properties, of bridging the gap between liquid and solid states. While most hydrogels are traditionally formed from polymers, biomolecules can also undergo gelation, as seen with proteins (e.g., collagen), enabling numerous applications. Peptide-based low-molecular-weight hydrogels (LMWHs), composed of amino acids, have emerged as innovative materials with a broad range of biomedical and biotechnological applications, gaining commercial interest in the 2010s. However, natural peptides composed solely of proteinogenic amino acids present several drawbacks, requiring structural or chemical modifications to enhance their performance. Additionally, multicomponent approaches, which involve combining multiple compounds to form hydrogels, have recently gained prominence as a promising strategy for developing more versatile and efficient systems. In this context, we explore emerging hybrid molecules, i.e., peptides functionalized with DNA bases (i.e., adenine, thymine, guanine, and cytosine), known as nucleopeptides. These compounds have shown encouraging results, yet much remains to be explored to unlock their full potential. In this study, we present a novel series of six (nucleo)-peptides derived from two distinct peptide sequences, Phe-Glu-Phe-Glu and Phe-Lys-Phe-Lys, negatively and positively charged at physiological pH, respectively, making them complementary in terms of electrostatic interactions. These peptides are functionalized with one of the four DNA nucleobases, introduced via a peptide nucleic acid (PNA) moiety. Thus, through a comprehensive multiscale systematic study, we report herein on the impact of charge complementarity and/or nucleobase-pair complementarity on the mechanical and physicochemical properties of the resulting multicomponent hydrogels (including gelation time, sol-gel transition temperature, stiffness, resistance to external stress, fibrillar network morphology, etc.). Then, the results highlight the undeniable potential of this approach, demonstrating that careful selection of components allows the fine-tuning of hydrogel properties. Interestingly, our findings reveal unexpected behaviors, underscoring the complexity of these bioinspired hybrid multicomponent systems while reinforcing their potential for the development of high-performance and innovative supramolecular hydrogels.
    Keywords:  DNA nucleobase; multicomponent hydrogels; multiscale analysis; nucleopeptide; rheology; supramolecular hydrogels; synergistic supramolecular assembly
    DOI:  https://doi.org/10.1021/acsmaterialsau.5c00240
  30. Nat Cell Biol. 2026 Jul 08.
      Nucleotides are essential for life, serving not only as the building blocks of the genome but also as cellular energy providers, metabolic cofactors and signalling molecules. To sustain cellular function and proliferation, cells must continuously generate, recycle and precisely balance nucleotide pools in response to fluctuating metabolic and environmental demands. Nucleotide metabolism is therefore not a static biosynthetic pathway, but a dynamic system tightly integrated with cell signalling and physiology. Here we highlight the regulatory logic of nucleotide metabolism, from acute post-translational regulation to transcriptional scaling, feedback control and higher-order spatial organization into multi-enzyme assemblies and filaments. Through the lens of human genetic disorders and cancer, we examine how nucleotide depletion, pool imbalance or intermediate toxicity produce striking tissue-selective pathologies. Together, these principles position nucleotide metabolism as a central regulatory axis linking cellular metabolism, signalling and fate in health and disease.
    DOI:  https://doi.org/10.1038/s41556-026-02004-9
  31. ACS Synth Biol. 2026 Jul 06.
      Bacteria can be engineered to produce double-stranded RNA (dsRNA) molecules that induce a targeted RNA interference (RNAi) response in plants and animals for applications ranging from pest control to functional genomics. We developed a genetically encoded sensor that uses bimolecular fluorescence complementation to report relative dsRNA levels within bacterial cells. We tested sensor designs consisting of fusions of different dsRNA-binding domains derived from viruses to fragments of a split fluorescent protein in Escherichia coli. Then, we used the optimized dsRNA sensor design to demonstrate enhanced dsRNA accumulation in engineered strains of the aphid symbiont Serratia symbiotica, including a new RNase III deletion mutant. Our biosensor provides a convenient fluorescent readout that can be used to accelerate the design-build-test cycle for maximizing dsRNA yields in bacteria, including species native to plant and animal microbiomes that can be used to implement symbiont-mediated RNAi.
    Keywords:  Genetically encoded biosensor; RNA interference; Serratia symbiotica; bimolecular fluorescence complementation; paratransgenesis; symbiont-mediated RNAi
    DOI:  https://doi.org/10.1021/acssynbio.6c00080
  32. ACS Synth Biol. 2026 Jul 09.
      Intracellular delivery of therapeutics remains a major challenge for modern medicine. To enhance intracellular uptake, therapeutics can be delivered with carrier proteins possessing an inherent cell-penetrating activity. There is an increasing need for new cell-penetrating carriers with diverse biophysical properties and mechanisms of action to transport a wide range of therapeutic cargo. As many cell-penetrating proteins and peptides derive from natural proteins, we sought to mine a previously unexplored community, the human gut microbiome, for cell-penetrating sequences. Here, we performed a high-throughput functional metagenomic screen to identify cell-penetrating protein fragments from the human gut microbiome. We identified protein fragments encoded within glycosidase enzymes from members of the Bacteroidetes phylum that mediate internalization into human cell lines when displayed on the surface of nonpathogenic, noninvasive Escherichia coli. We investigate one fragment, dubbed Gh_112, that adheres to human fibronectin, activates multiple endocytic pathways, and specifically promotes uptake of E. coli into multiple cancerous epithelial cell lines rather than healthy epithelial tissue in vitro. Overall, this work demonstrates that the human gut microbiome is a source of cell-penetrating sequences and expands the known repertoire of cell-penetrating carrier systems.
    Keywords:  bacterial display; cell-penetrating peptide; functional metagenomics; gut microbiome
    DOI:  https://doi.org/10.1021/acssynbio.6c00031
  33. Nat Commun. 2026 Jul 04.
      We learn to recognize a vast array of familiar objects, a process involving learning-related changes in inferotemporal cortex (IT) activity. A challenge to discovering mechanisms of familiarity learning is that it spans multiple timescales from minutes to days, and is accompanied by simultaneous changes in cellular, synaptic, and network properties. We leverage an integrated experimental-theoretical approach, using IT recordings in two male macaques during familiarity learning within and across sessions to infer underlying plasticity mechanisms. We identified two timescales of learning-related changes spanning minutes to days, consistent with distinct synaptic and cellular mechanisms. Across sessions, averaged responses gradually decreased with familiarity, consistent with synaptic plasticity. In contrast, within-session changes, including rapid response decay and increased spontaneous activity, aligned with intrinsic plasticity mechanisms. Recurrent networks endowed with learning rules inferred from experiments replicated the observed learning dynamics, supporting our hypothesis of distinct learning mechanisms - slow, synaptic plasticity at long timescales and fast, intrinsic plasticity at short timescales.
    DOI:  https://doi.org/10.1038/s41467-026-74791-6
  34. bioRxiv. 2026 Jun 29. pii: 2026.06.28.734975. [Epub ahead of print]
      Protein-protein interactions (PPIs) are central to biological processes. Designing small molecules that modulate dysregulated PPIs holds strong promise for targeting undruggable proteins. However, existing structure-based drug design approaches focus on well-defined small-molecule binding pockets and struggle to generalize to large, shallow, and chemically complex PPI interfaces. Here, we introduce Pep2Mol, a diffusion-based generative model for 3D molecule design that targets orthosteric PPI sites by explicitly incorporating binding peptides or proteins as structural guidance, moving beyond conventional pocket-conditioned generation. To enable model development and benchmarking, we curate a large-scale, high-quality dataset of 10,956 experimentally resolved protein complex structure pairs, each pairing an orthosteric competitive ligand with a protein binder at overlapping receptor interfaces. Pep2Mol integrates two SE(3)-equivariant graph neural networks that encode protein-ligand and protein-peptide interactions respectively, and fuses these representations via attention-based conditioning to jointly guide the diffusion trajectory. Extensive evaluations demonstrate that Pep2Mol generates chemically valid ligands with state-of-the-art binding affinities, providing a strong foundation for small-molecule inhibitor design against challenging PPI interfaces.
    DOI:  https://doi.org/10.64898/2026.06.28.734975
  35. ACS Appl Mater Interfaces. 2026 Jul 10.
      Electrochemiluminescence (ECL) has emerged as an ultrasensitive detection technique with zero background signal, and coupling of ECL with a commonly used fluorescence imaging technique can greatly advance the bioimaging/detection applications. However, molecular ECL luminophores have poor cellular uptake and cell-targeting properties. Here, we show that transforming a molecular ECL luminophore into a colloidal nanoprobe can increase its cellular uptake as well as enhance its application potential via both fluorescence and ECL. In particular, we have designed a colloidal magnetic nanoprobe that is composed of a 6-7 nm iron oxide core and surface terminated with a ruthenium complex that acts as both a fluorescent and ECL probe. The nanoparticles are functionalized with targeting ligands that enable the receptor-mediated uptake by specific cells. The designed nanoprobe has been used for imaging/detection of biotin/galactose overexpressed cells via both fluorescence and ECL, along with magnetic extraction of labeled cells for signal enhancement. Results demonstrate that molecular ECL luminophore can be transformed into a fluorescent-ECL nanoprobe with more advanced bioimaging and detection applications.
    Keywords:  [Ru(bpy)3]2+; bioimaging; cell targeting; electrochemiluminescence; fluorescence imaging; luminophores; nanoparticle
    DOI:  https://doi.org/10.1021/acsami.6c06590
  36. Nat Commun. 2026 Jul 06. pii: 5732. [Epub ahead of print]17(1):
      Supramolecular polymerization within living cells has emerged as a promising strategy for controlling cellular functions. Lipid droplets, intracellular organelles mainly composed of neutral lipids, provide low-polarity environments, yet their role in regulating supramolecular assembly pathways remains unclear. Here we show that triolein, a representative triacylglycerol featuring three ester groups and cis-9-octadecenyl chains, acts as an effective medium for kinetically controlled supramolecular polymerization. An alanine-based diamide-functionalized fluorophore forms supramolecular polymers in triolein through a nucleation-elongation mechanism, in which an initial nucleus triggers subsequent growth. Although the thermodynamic stability is comparable to that in di-n-butyl ether, a commonly used organic solvent, triolein suppresses spontaneous nucleation and inter-fiber bundling, creating a lag phase during which seeded polymerization guides monomers along a defined assembly pathway. Kinetic studies with ethyl oleate, a structural analogue of triolein, indicate that transient solute-solvent interactions contribute to the suppression of both nucleation and bundling events. This kinetic control enables stepwise seeded growth of multiblock nanostructures and establishes neutral lipids as functional media for the precision-controlled construction of supramolecular polymers.
    DOI:  https://doi.org/10.1038/s41467-026-74683-9
  37. Sci Adv. 2026 Jul 10. 12(28): eaee8616
      State-of-the-art soft materials can be engineered as sensors and actuators, yet, methods for learning from external information remain a subject of current research. Inspired by the use of large datasets to train artificial intelligence, tuning physical responsiveness to relayed data would introduce learning behavior in soft materials. In this work, we develop a trainable liquid crystal oligomer network (LCON) that stores digital information directly into its molecular configuration. By functionalizing the anisotropic LCON with photo-switchable azobenzene, we simultaneously integrate basic logic and memory in a material through a binary-state system; we coin this design the trainable self-propelled gate (T-SPG). We can tune the memory of our T-SPG with photonic stimuli, allowing the system to be trained by a conventional digital controller. We demonstrate the trainability of the T-SPG through two hierarchical tasks: a lower-level binary classification task where the decision boundary is stored as material memory, and a higher-level motion task that uses the stored memory to trigger actuation.
    DOI:  https://doi.org/10.1126/sciadv.aee8616
  38. Biomed Mater. 2026 Jul 08.
      Skin injuries, particularly severe injuries, such as full-thickness wounds or surgical sites, are susceptible to bacterial infection. Materials composed of keratin proteins have been investigated for use in wound dressings, and at least two products based on these materials have received FDA clearance for treating radiation dermatitis via the 510(k) mechanism. We hypothesized that hydrogels composed of oxidatively extracted keratins, known as keratose, could achieve sustained release of several classes of antibiotic drugs that might improve the performance of keratins as wound dressing materials. Reverse-phase HPLC methods were developed to quantify release of ciprofloxacin, cefazolin, and neomycin antibiotics from keratose hydrogels. Nearly 100% of ciprofloxacin and cefazolin were released from the hydrogels within 7 days, while only approximately 20% of neomycin was released. In a combination keratose hydrogel containing neomycin and cefazolin, the amount of cefazolin release decreased (compared to hydrogels with only cefazolin) to about 60% over 7 days whereas neomycin release increased (compared to hydrogels with only neomycin) to nearly 30%. The antibiotics released from the hydrogels inhibited growth of Gram-positive (Staphylococcus aureus) and Gram-negative (Pseudomonas aeruginosa) bacteria in an in vitro broth inhibition assay. Keratose hydrogels containing a combination of the first-line antibiotics neomycin and cefazolin inhibited S. aureus for 18 days and P. aeruginosa for 6 days which was comparable to or better than hydrogels containing broad-spectrum ciprofloxacin that had effect for 11 days. This work suggests that modification of keratin biomaterials with antibiotics may enhance their utility by inhibiting bacterial infections.
    Keywords:  cefazolin; ciprofloxacin; high-performance liquid chromatography; keratin; keratose; neomycin
    DOI:  https://doi.org/10.1088/1748-605X/ae8817
  39. Science. 2026 Jul 09. eaed6782
      Phages often degrade the genome of their bacterial host to individual nucleotides. Here we describe Metis, a bacterial defense system that directly senses phage-mediated host genome degradation. Metis aborts phage infection once it detects the modified mono-nucleotide m6dAMP. As methylation of deoxyadenosines usually occurs on the DNA polymer, accumulation of m6dAMP signals that the host genome has been degraded. In type I Metis, sensing of m6dAMP activates an NAD+ diphosphatase, leading to NAD+ depletion and cessation of the infection process; while the effector in type II Metis is a membrane-spanning protein whose toxicity is triggered in response to the modified mono-nucleotide. We further show that Metis defense depends on endogenous DNA methylases, and that phages can escape Metis via mutations that inactivate host genome degradation.
    DOI:  https://doi.org/10.1126/science.aed6782
  40. Phys Rev Lett. 2026 Jun 19. 136(24): 248402
      Fracture typically signifies mechanical failure in engineering materials, whereas controlled cracking may actively sculpt tissues through precise biological regulation. Here, we establish a multiscale nonlinear peridynamic theory that accounts for cellular mechanosensing to decipher the spontaneous fracture of active tissues. We show that tissues cultured in a ring-shaped domain can undergo periodic fracture to generate multicellular aggregates with regular spacing, recapitulating prior morphogenetic experiments on avian dermal cell collectives. It is found that the number of cracks varies nonmonotonically with the substrate stiffness. We predict that a narrow tissue favors equally spaced radial cracking, while such ordered cracks deflect, branch, and randomize increasingly as the tissue broadens, attributable to the anisotropy-isotropy transition of tissue stresses induced by the interplay of active contraction and domain geometry. Backed by energetic arguments, we identify the factors that control the characteristic size of tissue fracture. Our Letter reveals a synergy of physics, geometry, and cellular mechanosensing in controlling active tissue fracture to achieve tissue-level organization.
    DOI:  https://doi.org/10.1103/hw7t-kf32
  41. Nature. 2026 Jul 08.
    Tabula Sapiens Consortium
      Developing a universal representation space for cells that encompasses the tremendous molecular diversity of cell types across species would be transformative for cell biology. Recent work using single-cell transcriptomic approaches to create molecular definitions of cell types in the form of cell atlases has provided the necessary data for such an endeavour1-3. Here we present the universal cell embedding (UCE) foundation model. UCE was trained on a large corpus of cell data using self-supervision, creating a unified biological latent space that can represent cells across diverse tissues and species. This latent space captures important biological variation despite the presence of experimental noise. UCE's universality means that new cells can be embedded with no data labelling, model training or fine-tuning. We used UCE to create the Integrated Mega-scale Atlas, embedding 36 million cells, with more than 1,000 uniquely named cell types, from hundreds of experiments, dozens of tissues and eight species. We gain insights into the organization of cell types and tissues within the space. UCE's embedding space exhibits emergent behaviour, identifying biology that it was never trained for, such as identifying developmental lineages and embedding data from species that were not included in the training set. Overall, by enabling a universal representation for every cell state and type, UCE is a valuable tool for analysis, annotation and hypothesis generation over single-cell data.
    DOI:  https://doi.org/10.1038/s41586-026-10689-z