bims-cebooc Biomed News
on Cell biology of oocytes
Issue of 2026–03–15
eleven papers selected by
Gabriele Zaffagnini, Universität zu Köln



  1. Nat Commun. 2026 Mar 11.
      Oocyte development requires tight regulation of transcription and RNA metabolism, which is coordinated by RNA-binding proteins, whose roles in mammalian oogenesis remain incompletely understood. Here, we identify the DEAD-box RNA helicase DDX5 as a key regulator of RNA homeostasis in oocytes. Oocyte-specific deletion of DDX5 leads to female sterility, which is characterized by defective chromatin remodeling, meiotic arrest, increased aneuploidy, and fertilization failure. Mechanistically, DDX5 maintains RNA homeostasis through three interconnected processes: (1) promoting transcription via interaction with RNA polymerase II in nonsurrounded nucleolus-stage germinal vesicle oocytes; (2) clearing retrotransposon RNAs to safeguard transcriptome integrity; and (3) supporting maternal mRNA storage by coordinating nuclear export, mitochondrial organization, and mitochondria-associated ribonucleoprotein domain assembly. Our study establishes DDX5 as a master regulator that integrates transcriptional and post-transcriptional programs to ensure oocyte competence and fertility.
    DOI:  https://doi.org/10.1038/s41467-026-70237-1
  2. Elife. 2026 Mar 11. pii: RP109358. [Epub ahead of print]14
      Mouse female primordial germ cells (PGCs) undergo five synchronous, incomplete mitotic divisions and send each resulting germline cyst into meiosis to fragment and produce 4-6 oocytes and 24-26 supportive nurse cells. However, no system of polarity has been found to specify mammalian oocytes, link them appropriately to nurse cells and enable them to acquire high-quality organelles and cytoplasm. We report that mouse cysts develop an asymmetric Golgi, endoplasmic reticulum (ER), and microtubule-associated 'fusome,' similar to the oocyte-determining fusome in Drosophila cysts. The mouse fusome distributes asymmetrically among cyst cells and enriches in future oocytes with Pard3 and Golgi-endosomal UPR (unfolded protein response) proteins. Spindle remnants rich in stable acetylated microtubules, like those building the Drosophila and Xenopus fusomes, transiently link early mouse cyst cells for part of each cell cycle. A non-random gap in these microtubules predicts that initial cysts fragment into similar six-cell derivatives, providing a potential mechanism for producing uniform oocytes. Together with previous studies, these results argue that a polarized fusome underlies the development of female gametes from the PGC to follicular oocyte stages in diverse animals including mammals.
    Keywords:  apical-basal polarity; cell cycle; developmental biology; fusome; germline cyst; mouse; oocyte development
    DOI:  https://doi.org/10.7554/eLife.109358
  3. Development. 2026 Mar 12. pii: dev.205203. [Epub ahead of print]
      Female reproductive senescence results from the regulated depletion of a finite pool of oocytes called the ovarian reserve. This pool of oocytes is initially established during fetal development, but the oocytes that comprise it must remain quiescent for decades until they are activated during maturation in adulthood. In order for developmentally competent oocytes to populate the ovarian reserve they must successfully initiate both meiosis and oogenesis. As the factors that regulate the timing and fidelity of these early events remain elusive, we assessed the precise function and timing of the transcriptional regulator TAF4b during meiotic prophase I progression in mouse fetal oocytes. Compared to matched controls, E14.5 Taf4b-deficient oocytes enter meiosis I in a timely manner; however, their subsequent progression through the pachytene-to-diplotene transition of meiotic prophase I is compromised. Moreover, this disruption of meiotic progression is associated with the reduced ability of Taf4b-deficient oocytes to repair double-strand DNA breaks. Transcriptional profiling of Taf4b-deficient oocytes reveals that between E16.5 and E18.5 these oocytes fail to properly coordinate the reduction of meiotic gene expression and the activation of oocyte differentiation genes.
    Keywords:  Female infertility; Fetal oocyte attrition; Meiosis; Oogenesis; Ovarian reserve; Transcription
    DOI:  https://doi.org/10.1242/dev.205203
  4. Sci Adv. 2026 Mar 13. 12(11): eaeb2345
      The spindle assembly checkpoint (SAC) is vital for preventing oocyte aneuploidy, a leading cause of female infertility, miscarriages, and trisomy syndromes. However, whether deubiquitination participates in SAC regulation remains unknown. Here, we reveal that the deubiquitinase USP8 acts as a SAC regulator to prevent aneuploid egg formation. Mechanistically, depletion of USP8 inactivates the SAC, accelerates meiotic progression, and causes abnormal spindle assembly and chromosome alignment, ultimately leading to aneuploidy. Intriguingly, we identify USP8 in oocytes as a previously unidentified interaction partner of BUB3, a key component of the SAC, and demonstrate that USP8 stabilizes BUB3 through its deubiquitinating activity. Moreover, exogenous BUB3 rescues the defects observed in USP8-depleted oocytes. Together, our findings not only clarify that deubiquitination participates in regulating the SAC in oocytes but also uncover a unique role for USP8 in controlling the SAC via its interaction with BUB3.
    DOI:  https://doi.org/10.1126/sciadv.aeb2345
  5. Reproduction. 2026 Mar 12. pii: xaag031. [Epub ahead of print]
      Farnesoid X receptor (FXR), a ligand-dependent nuclear receptor predominantly expressed in enterohepatic tissues, is also present in the ovary; however, its role in female reproduction remains unclear. In this study, we investigated the role of FXR in regulating the ovarian reserve using a novel FXR knockout (FXR-KO) mouse. FXR-KO mice exhibited an increased number of ovulated oocytes, associated with an enlarged pool of secondary follicles, suggesting enhanced recruitment from the primordial follicle pool. During the neonatal period, FXR-KO ovaries showed accelerated primordial follicle activation, evidenced by increased Forkhead box O3 (FOXO3) nuclear exclusion, while pharmacological activation of FXR suppressed this process in wild-type ovaries. Mechanistically, FXR directly bound to the FoxO3 promoter, increasing its expression without altering FOXO3 phosphorylation, indicating transcriptional regulation. In vitro, activation of FXR in granulosa cells by an agonist upregulated cell cycle inhibitors and suppressed proliferation, suggesting FXR maintains primordial follicle dormancy by restraining granulosa cell differentiation and potentially enhancing oocyte quiescence. These findings identify FXR as a novel regulator of ovarian reserve maintenance, highlighting its potential as a pharmacological target to modulate reproductive lifespan and fertility management in mammals.
    Keywords:  Cell cycle control; Follicle; Gene regulation; Granulosa cells; Ovary
    DOI:  https://doi.org/10.1093/reprod/xaag031
  6. Biol Reprod. 2026 Mar 13. pii: ioag061. [Epub ahead of print]
      KIT signaling is a fundamental regulatory pathway that preserves cellular homeostasis and controls cell development and fate across a wide range of organs and cell types. Consistent with this pleiotropic role, mutations in c-KIT/Kit have been associated with a wide range of phenotypes, including sterility, piebaldism, nevus formation, mastocytosis, and multiple malignancies. The contribution of c-KIT/Kit to reproductive function has attracted sustained attention for several decades, underscoring its essential role in fertility and gonadal biology. KIT expression is observed in oocytes - localized to the oocyte membrane and the cytoplasm - as well as in theca cells and interstitial cells, suggesting a multifaceted role in follicular development. Notably, all Kit mutant models develop primary ovarian insufficiency (POI) with variable onset, characterized by endocrine dysfunction, impaired folliculogenesis, and eventual female infertility. These findings collectively establish KIT signaling as a critical regulator of ovarian integrity, as both gain- or loss-of-function mutations in Kit consistently recapitulate POI-associated phenotypes. However, despite substantial progress, the precise molecular mechanisms by which KIT signaling integrates these pathways to preserve primordial follicle survival and prevent POI remain incompletely understood. Here, we summarize current knowledge of KIT expression and the functional consequences of Kit mutations, with particular emphasis on oocytes across ovarian cell populations and in comparison to other organ systems in humans and mice. We further evaluate the physiological and pathological significance of ovarian KIT signaling in female fertility and highlight crucial knowledge gaps that must be addressed to fully elucidate its role in maintaining ovarian function.
    Keywords:  KIT; KIT ligand; folliculogenesis; other tissues; ovary
    DOI:  https://doi.org/10.1093/biolre/ioag061
  7. Fertil Steril. 2026 Mar 11. pii: S0015-0282(26)00156-1. [Epub ahead of print]
       OBJECTIVE: To investigate spindle dynamics and chromosome segregation in human preimplantation embryos. Chromosome segregation errors that cause aneuploidy are frequent in human preimplantation development. However, direct studies of the mechanisms of cell division in human embryos to understand the root causes of these errors are few.
    DESIGN: Preimplantation human embryos were stained with SPY650-DNA and SPY555-tubulin to visualize mitotic chromosomes and microtubules, respectively, and spindle dynamics and chromosome segregation were analyzed using confocal live imaging.
    SUBJECTS: 91 cryopreserved embryos donated by 26 patients aged 24-41 years old (mean age 34 years).
    EXPOSURE: We used live imaging to analyze spindle assembly and chromosome segregation in human preimplantation embryos donated to research.
    MAIN OUTCOME MEASURE(S): Mitosis duration, microtubule dynamics, mitotic defects.
    RESULTS: We find that spindle assembly occurs in an 'outside-in' manner, followed by a canonical anaphase chromosome segregation, and a persistent spindle remnant that connects sister cells for several hours after anaphase. We find that chromosome segregation errors occur both at Day 2/3 and also in Day 5 embryos, including the generation of micronuclei that undergo a non-canonical inheritance pattern that is likely a major contributor to blastocyst aneuploidy.
    CONCLUSIONS: Our data provide a foundational understanding of chromosome segregation mechanisms for further unravelling of the causes of segregation error, and highlight micronuclei as a central player in aneuploidy genesis in human embryos.
    Keywords:  aneuploidy; embryo; microtubule; mitosis
    DOI:  https://doi.org/10.1016/j.fertnstert.2026.03.004
  8. Commun Biol. 2026 Mar 13.
      Age-dependent reproductive decline has become a significant global health concern as the average maternal age at first birth increases. Fertility loss associated with reproductive aging is driven in part by alterations to ovarian composition and function, dysregulation of folliculogenesis, and increased inflammatory signaling. Our understanding of the molecular changes underlying ovarian aging has been expanded by single-cell and spatial transcriptomic studies, which identified infiltration of immune cells as a feature of ovarian aging. However, the function of these age-associated immune cells and their potential contributions to the inflammaging phenotype remain unclear. In this study, we integrate single-cell and spatial transcriptomics to define changes in the composition and intercellular signaling in the aging mouse ovary. We identify specific macrophage and T cell subpopulations that increase with age and are key sources of pro-inflammatory signaling in old ovaries. Further, we predict bidirectional signaling between these pro-inflammatory cells and granulosa cell populations that may impair follicular growth and development while promoting immune cell recruitment. These findings provide insights into the mechanisms that drive ovarian inflammaging.
    DOI:  https://doi.org/10.1038/s42003-026-09826-1
  9. Dev Biol. 2026 Mar 11. pii: S0012-1606(26)00058-8. [Epub ahead of print]
      Tissue homeostasis is dependent on precise coordination between endocrine organs in response to changes in organism physiology. Secreted circulating factors from adipocytes regulate the behavior of stem cell lineages in peripheral tissues in multiple organisms. In addition to their endocrine roles, Drosophila adipocytes store and secrete amino acid storage proteins throughout development. During the larval feeding period, adipocytes secrete storage proteins into the hemolymph, which are reabsorbed by the adipose tissue during metamorphosis to control adult organ size and fertility. Despite the known functions for storage proteins during the larval stages, their requirement during Drosophila adulthood and reproduction are uncharacterized. We discover that adipocyte-specific knockdown of the storage proteins Larval serum protein 1 (Lsp1) α/β/γ and Larval serum protein 2 (Lsp2) results in a decrease in GSC maintenance. We further reveal that decreased GSC number is due to downregulation of Target of Rapamycin (TOR) signaling in GSCs, suggesting compromised amino acid sensing directly in GSCs. We also find that the proteins that mediate storage protein adipocyte reabsorption, Fat body protein 1 (Fbp1) and Fat body protein 2 (Fbp2), are expressed in ovarian follicle cells. Intriguingly, Fbp1 nor Fbp2 appear to be required in follicle cells for GSC maintenance, suggesting undiscovered requirements for amino acid storage proteins in oogenesis. Our results highlight a novel role for Drosophila amino acid storage proteins during adulthood and in regulating tissue stem cell lineages.
    Keywords:  Fbp1; Fbp2; Lsp1; Lsp2; Oogenesis
    DOI:  https://doi.org/10.1016/j.ydbio.2026.03.004
  10. Nat Commun. 2026 Mar 11.
      The piwi-interacting RNAs (piRNAs) are essential for controlling transposable elements (TEs) activity in germ cells, yet their expression dynamics and functions during human oogenesis remain poorly understood. Here, we simultaneously profile small and long RNAs in individual human oocytes across four developmental stages. piRNAs, especially PIWIL3-associated short piRNAs, are the predominant small non-coding RNAs during human oogenesis. A marked increase in short-piRNAs coincide with a global reduction of TE expression, particularly LINE-1 and endogenous retroviruses (ERVs). In contrast, PIWIL1- and PIWIL2-associated long piRNAs correlate with the downregulation of certain specific ERV subfamilies. Genomic analyses reveal that highly productive piRNA clusters have evolved asymmetric antisense insertion bias toward LINE-1 and ERVs, enabling TE families-specific regulation. Together, our study provides a valuable single-cell dataset of small and long RNA co-expression landscapes in developing human oocytes and reveals coordinated yet distinct roles of different PIWI/piRNA classes in TE repression during human oogenesis, with short-piRNAs acting as the primary and broad-spectrum suppressors, and long-piRNAs providing coordinated ERV-specific repression.
    DOI:  https://doi.org/10.1038/s41467-026-70296-4
  11. Curr Biol. 2026 Mar 09. pii: S0960-9822(26)00128-4. [Epub ahead of print]36(5): R210-R228
      Aneuploidy, an imbalance in the number of chromosomes, is highly detrimental at both the cellular and the organismal levels and is the major cause of miscarriages in humans. Despite its deleterious effects, recent data have revealed a high prevalence of aneuploidy in early human embryos. On one hand, organismal aneuploidy is strikingly common due to the high frequency of aneuploid gametes, particularly aged oocytes. On the other hand, mosaic aneuploidy has been recently shown to be also widespread in early embryos, due to a high rate of chromosome segregation errors during the first post-zygotic divisions. Of note, aneuploid mosaic embryos can result in healthy individuals. Here, we review the compatibility of aneuploidy with correct embryo development and also address the presence of aneuploidy in normal somatic tissues, and the implication of aneuploid mosaicism for the processes of aging and tumorigenesis. We place particular emphasis on how different kinds of aneuploidies, either trisomies or monosomies, whole-chromosome or segmental, differentially contribute to the discussed processes. Finally, we highlight how cell competition may serve as a crucial mechanism by which both developing and somatic tissues deal with the emergence of aneuploidy.
    DOI:  https://doi.org/10.1016/j.cub.2026.01.056